biology phd schools

Ph.D. Program

The training for a Ph.D. in Biology is focused on helping students achieve their goals of being a successful research scientist and teacher, at the highest level. Students work closely with an established advisor and meet regularly with a committee of faculty members to facilitate their progress. The Biology Ph.D. program is part of the larger Biosciences community at Stanford, which includes doctorate programs in the basic science departments at Stanford Medical School. 

There are two tracks within the Biology Ph.D. program:

• Cell, Molecular and Organismal Biology
• Ecology and Evolution

(Previously a part of the Department of Biology Hopkins Marine Station is now a part of the Oceans Department within  Stanford Doerr School of Sustainability )

All  tracks are focused on excellence in research and teaching in their respective areas; where there are differences between the tracks, they are indicated in the links below. 

Requirements & Forms

Dissertation defense, cellular and molecular biology training program, stanford biology ph.d. preview program, career development resources.

Biology Graduate Programs in America

1-25 of 396 results

MIT School of Science

Cambridge, MA •

Massachusetts Institute of Technology •

Graduate School

Massachusetts Institute of Technology ,

Graduate School ,

CAMBRIDGE, MA ,

Stanford University Department of Humanities and Sciences

Stanford, CA •

Stanford University •

Stanford University ,

STANFORD, CA ,

Stanford University School of Medicine

• • Rating 5 out of 5   3 reviews

Doctoral Student: I begin in the Fall of 2023--Based on my interview, and pre-classes I have taken this program is amazing. It is very student-centered. ... Read 3 reviews

3 Niche users give it an average review of 5 stars.

Featured Review: Doctoral Student says I begin in the Fall of 2023--Based on my interview, and pre-classes I have taken this program is amazing. It is very student-centered. .

Read 3 reviews.

Shippensburg University of Pennsylvania

Graduate School •

SHIPPENSBURG, PA

• • Rating 4.85 out of 5   13

College of Science and Engineering - San Francisco State University

San Francisco State University •

SAN FRANCISCO, CA

College of Liberal Arts and Social Sciences - University of Illinois at Springfield

University of Illinois Springfield •

SPRINGFIELD, IL

Harvard Graduate School of Arts and Sciences

Harvard University •

• • Rating 4.56 out of 5   9 reviews

Other: I am Harvard Extension School student pursuing a master degree, ALM, in sustainability. I have achieved a 3.89 in this program so far and have qualified, applied, and accepted as a 'Special Student' in the Harvard Graduate School of Arts and Sciences. Through this School, I will be focusing my time at the John A. Paulson school of Engineering & Applied Sciences. Looking forward to wrapping up my final year on campus! ... Read 9 reviews

Harvard University ,

9 Niche users give it an average review of 4.6 stars.

Featured Review: Other says I am Harvard Extension School student pursuing a master degree, ALM, in sustainability. I have achieved a 3.89 in this program so far and have qualified, applied, and accepted as a 'Special Student'... .

Read 9 reviews.

Harvard Medical School

Boston, MA •

• • Rating 4.94 out of 5   16 reviews

Other: Harvard is my dream school. Next year I will graduate the Mongolian National University of Medical Sciences. Then I will applying to Harvard Medical School. If I accepted by this school, I will try my best of the best….. ... Read 16 reviews

BOSTON, MA ,

16 Niche users give it an average review of 4.9 stars.

Featured Review: Other says Harvard is my dream school. Next year I will graduate the Mongolian National University of Medical Sciences. Then I will applying to Harvard Medical School. If I accepted by this school, I will try... .

Read 16 reviews.

Duke University School of Medicine

Durham, NC •

Duke University •

• • Rating 4.76 out of 5   17 reviews

Doctoral Student: Professors are incredible, always willing to go the extra mile to help students succeed. Curriculum is designed to prepare students to be successful therapists. This program is truly incredible. ... Read 17 reviews

Duke University ,

DURHAM, NC ,

17 Niche users give it an average review of 4.8 stars.

Featured Review: Doctoral Student says Professors are incredible, always willing to go the extra mile to help students succeed. Curriculum is designed to prepare students to be successful therapists. This program is truly incredible. .

Read 17 reviews.

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Brown University Graduate School

Providence, RI •

Brown University •

Brown University ,

PROVIDENCE, RI ,

School of Arts & Sciences - University of Pennsylvania

Philadelphia, PA •

University of Pennsylvania •

University of Pennsylvania ,

PHILADELPHIA, PA ,

Weinberg College of Arts and Sciences

Evanston, IL •

Northwestern University •

Northwestern University ,

EVANSTON, IL ,

Vanderbilt University School of Medicine

Nashville, TN •

Vanderbilt University •

• • Rating 5 out of 5   2 reviews

Graduate Student: Vanderbilt University School of Nursing. Loved my experience! Great teachers, easy to access, clear cut goals in class, great clinical experiences. ... Read 2 reviews

Vanderbilt University ,

NASHVILLE, TN ,

2 Niche users give it an average review of 5 stars.

Featured Review: Graduate Student says Vanderbilt University School of Nursing. Loved my experience! Great teachers, easy to access, clear cut goals in class, great clinical experiences. .

Read 2 reviews.

Washington University in St. Louis - Arts & Sciences

St. Louis, MO •

Washington University in St. Louis •

Washington University in St. Louis ,

ST. LOUIS, MO ,

Guarini School of Graduate and Advanced Studies

Hanover, NH •

Dartmouth College •

• • Rating 5 out of 5   1 review

Alum: I had a beautiful life-changing experience at the grand Guarini School of Graduate and Advanced Studies. The Guarini graduate program MALS created so many beautiful opportunities in interdisciplinary learning. Guarini is well integrated within the College, which provides a unique world-class learning experience. Guarini went beyond my expectations and made this experience the best I had and will ever have in my life. The only thing I would like changed is having a GRAD diploma in Latin instead of English to keep up with College tradition. ... Read 1 review

Dartmouth College ,

HANOVER, NH ,

1 Niche users give it an average review of 5 stars.

Featured Review: Alum says I had a beautiful life-changing experience at the grand Guarini School of Graduate and Advanced Studies. The Guarini graduate program MALS created so many beautiful opportunities in interdisciplinary... .

Read 1 reviews.

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Division of Biology and Biological Engineering - California Institute of Technology

Pasadena, CA •

California Institute of Technology •

California Institute of Technology ,

PASADENA, CA ,

College of Science - University of Notre Dame

Notre Dame, IN •

University of Notre Dame •

University of Notre Dame ,

NOTRE DAME, IN ,

Biological Sciences Division - University of Chicago

Chicago, IL •

University of Chicago •

• • Rating 4 out of 5   1 review

Master's Student: So far so good. Nothing out of the ordinary. Professors are knowledgeable. I would say to ensure that you know your interests and work closely with the advisors to pick suitable classes. ... Read 1 review

University of Chicago ,

CHICAGO, IL ,

1 Niche users give it an average review of 4 stars.

Featured Review: Master's Student says So far so good. Nothing out of the ordinary. Professors are knowledgeable. I would say to ensure that you know your interests and work closely with the advisors to pick suitable classes. .

Pritzker School of Medicine

Dornsife college of letters, arts and sciences.

Los Angeles, CA •

University of Southern California •

University of Southern California ,

LOS ANGELES, CA ,

Cornell University College of Arts & Sciences

Ithaca, NY •

Cornell University •

Cornell University ,

ITHACA, NY ,

Georgetown University School of Medicine

Nw Washington, DC •

Georgetown University •

• • Rating 4.75 out of 5   4 reviews

Doctoral Student: Compressed pre-clinical work. Administration has been working on finding better and better educators, as well as hiring more learning specialists for the student body. Could have more focus on Step 1 and Step 2 preparation. ... Read 4 reviews

Georgetown University ,

NW WASHINGTON, DC ,

4 Niche users give it an average review of 4.8 stars.

Featured Review: Doctoral Student says Compressed pre-clinical work. Administration has been working on finding better and better educators, as well as hiring more learning specialists for the student body. Could have more focus on Step 1... .

Read 4 reviews.

School of Arts and Sciences - Tufts University

Medford, MA •

Tufts University •

• • Rating 4 out of 5   3 reviews

Master's Student: Best: - Incredible colleagues, many of whom are working professionals in the field, all of whom are passionate & come from diverse backgrounds & bring a wide variety of life/ professional experiences that enhance the overall experience -One of the few DEIJ degree programs offered worldwide -flexible options for program completion -many passionate professors & deep learning opportunities -increasing program investment by Tufts, including 2 new FT program-specific professors Most challenging: -program has grown exponentially in the last several years & has experienced commiserate growing pains, including confusion about requirements, need for more faculty, and more opportunities for practice. Tufts is responding but needs to continue its investment. -Since the program is interdisciplinary, students in the DEIJ-L program often need to take classes offered by other dept's/ colleges which may have different requirements & DEIJ-L students may not get preference for registration purposes ... Read 3 reviews

Tufts University ,

MEDFORD, MA ,

3 Niche users give it an average review of 4 stars.

Featured Review: Master's Student says Best: - Incredible colleagues, many of whom are working professionals in the field, all of whom are passionate & come from diverse backgrounds & bring a wide variety of life/ professional experiences that... -One of the few DEIJ degree programs offered worldwide -flexible options for program completion -many passionate professors & deep learning opportunities -increasing program investment by Tufts, including 2 new FT program-specific professors Most challenging: -program has grown exponentially in the last several years & has experienced commiserate growing pains, including confusion about requirements, need for more faculty, and more opportunities for... -Since the program is interdisciplinary, students in the DEIJ-L program often need to take classes offered by other dept's/ colleges which may have different requirements & DEIJ-L students may not... .

UCLA College of Letters and Science

University of California - Los Angeles •

• • Rating 3 out of 5   1 review

University of California - Los Angeles ,

1 Niche users give it an average review of 3 stars.

Krieger School of Arts & Sciences

Baltimore, MD •

Johns Hopkins University •

• • Rating 4.53 out of 5   19 reviews

Master's Student: I have yet to enroll for Fall 2023 after receiving my acceptance letter due to a delay in my need-based financial aid from JHU. However the Homewood Campus in Baltimore is beautiful and my Student Advisor, Alexis has been extremely helpful in initiating my enrollment process and answering all of my questions in a timely matter. My intended Advanced Academic Program is the accelerated (2 semester), dual-modality, 40-credit M.S. in Biotechnology, Biodefense concentration. All of the anticipated course subjects are diverse and there's even a customizable core lab course on campus (at least until Summer 2024). I can't wait and I wish you all the best in your search for academic programs or professional certifications. ... Read 19 reviews

Johns Hopkins University ,

BALTIMORE, MD ,

19 Niche users give it an average review of 4.5 stars.

Featured Review: Master's Student says I have yet to enroll for Fall 2023 after receiving my acceptance letter due to a delay in my need-based financial aid from JHU. However the Homewood Campus in Baltimore is beautiful and my Student... .

Read 19 reviews.

Mellon College of Science

Pittsburgh, PA •

Carnegie Mellon University •

Blue checkmark.

Carnegie Mellon University ,

PITTSBURGH, PA ,

The Graduate School of Arts & Sciences - University of Virginia

Charlottesville, VA •

University of Virginia •

Alum: Very good in some areas, excellent in other areas, many academic choices available in all areas of study ... Read 1 review

University of Virginia ,

CHARLOTTESVILLE, VA ,

Featured Review: Alum says Very good in some areas, excellent in other areas, many academic choices available in all areas of study .

College of Sciences - Georgia Institute of Technology

Atlanta, GA •

Georgia Institute of Technology •

Other: Georgia Tech is among the top colleges you can attend. My journey there had its highs and lows, but looking back, every experience contributed significantly to my growth. A piece of advice I'd offer to newcomers is to prioritize building friendships early on. One regret I have as I reflect on my year is not connecting with more people. I initially thought I should always work independently, but that was a misstep in my academic journey. Remember, it's essential to strike a balance between learning and having fun. ... Read 1 review

Georgia Institute of Technology ,

ATLANTA, GA ,

Featured Review: Other says Georgia Tech is among the top colleges you can attend. My journey there had its highs and lows, but looking back, every experience contributed significantly to my growth. A piece of advice I'd offer... .

College of Arts and Sciences - American University

American University •

WASHINGTON, DC

Lehigh University

BETHLEHEM, PA

• • Rating 4.42 out of 5   19

University of Illinois Springfield

• • Rating 4.29 out of 5   31

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Biology, PhD

The Biology Graduate Program represents many areas of biology, and interactions with a diverse group of colleagues provide opportunities to broaden every student’s thinking and make connections between different fields and scientific approaches. Areas of research include microbiology, cell biology, development, physiology, neuroscience, animal behavior, plant biology, genetics, computational biology, evolution, ecology and biodiversity. 

Each entering graduate student has the freedom to pursue topics ranging from the behavior of molecules to that of cells, organisms, genomes, and ecosystems. We encourage students to get broad exposure through lab rotations with any faculty member in the Biology Graduate Group. As students focus on more specific research interests, they tailor their graduate education accordingly, choosing courses from different departments and schools at Penn as appropriate.

Students complete most of their course work and lab rotations in the first year and then start their thesis research in the second year while completing their teaching requirement and preparing for their candidacy exams.  Students are then fully focused on thesis research by the end of the second year.  Students still have the option of taking additional courses in advanced years in order to enhance their graduate research.

For more information: http://www.bio.upenn.edu/graduate/

View the University’s Academic Rules for PhD Programs .

Required Courses

The total course units for graduation in this program is 13.5.

See the website for a list of electives:  http://www.bio.upenn.edu/graduate/handbook/academic-topics/course-requirements

The degree and major requirements displayed are intended as a guide for students entering in the Fall of 2023 and later. Students should consult with their academic program regarding final certifications and requirements for graduation.

Sample Plan of Study

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Doctor of Philosophy (PhD) in Biology

The PhD in Biology is a research degree requiring graduate-level coursework, completion of a dissertation, and two semesters of participation in teaching (usually as a teaching fellow in laboratory or discussion sections of lecture courses led by Biology faculty). For most students, obtaining this degree typically involves five or more years of full-time study.

A summary of Biology PhD student expectations by year can be found here . Full details can be found in the Graduate Program Guide .

The Biology Department guarantees support for five years for all PhD students, contingent on satisfactory performance in the program.

How to Apply    Frequently Asked Questions

Learning Outcomes

1. Demonstrate academic mastery in one of three areas of Biology: Ecology, Evolution, Behavior & Marine Biology; Neurobiology; or Cellular & Molecular Biology.

2. Attain research expertise , including grant writing experience, and complete original research that advances a specific field of study within one of three broad subject areas represented in the department: Ecology, Behavior, Evolution & Marine Biology; Neurobiology; or Cellular & Molecular Biology.

3. Attain teaching experience and expertise in one of three broad areas of Biology: Ecology, Behavior, Evolution & Marine Biology; Neurobiology; or Cellular & Molecular Biology.

4. Attain the skills and qualifications needed for employment in an academic, government, or private sector position related to the life sciences.

Coursework Requirements

Students must complete 64 credits with a minimum grade point average of 3.0; at least 32 of these credits must be accrued from lecture, laboratory, or seminar courses. Students with prior graduate work may be able to transfer course credits.  See the  Graduate School of Arts & Sciences (GRS) Transfer of Credits policy  for more details. 

• 2 semesters of Progress in Research Seminars (2 credits each):

Cell & Molecular Biology:  BI 583 & BI 584 (CM section)

Ecology, Behavior, Evolution & Marine Biology:  BI 579 & BI 580

Neurobiology:   BI 583 & BI 584 (NEURO section)

• 1 semester grant-writing course (2 credits):

Cell & Molecular Biology:  BI 581

Ecology, Behavior, Evolution & Marine Biology: BI 671

Neurobiology: BI 581

• 1 semester pedagogy course (1 credit): All Biology PhD students take  BI 697
• 1 quantitative course (3–4 credits): from a list of recommended courses .
• Additional courses vary by specific track below:

Cell & Molecular Biology

1. GRS BI 791/GRS BI 792 Graduate Rotation Credits (2 credits each/4 credits total)

2. GRS BI 753 Advanced Molecular Biology  (4 credits)

3. GRS MB 721 Graduate Biochemistry  (4 credits)

4. GRS BI 735 Advanced Cell Biology  (4 credits)

5. Two electives (8 credits, 500-level and above), see the Graduate Program Guide for recommendations

6. Research credits  (remaining credits)

Ecology, Behavior, Evolution & Marine Biology

1. Six electives  (24 credits, 500-level and above), see the Graduate Program Guide for recommendations

2. Research credits  (remaining credits)

Ecology, Behavior, Evolution & Marine Biology PhD candidate coursework is highly variable. Students, in consultation with advisors, develop a plan of coursework and research. Students are required to take a minimum of 32 credits of coursework. The remainder of the credits should be research.

Neurobiology

2. GRS BI 755 Cellular and Systems Neuroscience  (4 credits)

3. GRS BI 741 Neural Systems: Functional Circuit Analysis  (4 credits)

4. Four electives  (16 credits, 500-level and above), see the Graduate Program Guide for recommendations

5. Research credits  (remaining credits)

Teaching Requirement

The department requires a minimum of two semesters of teaching as part of the Doctor of Philosophy program. During the first semester of teaching, students are required to enroll in our first-year seminar course, GRS BI 697 A Bridge to Knowledge . The course provides guidance and training on pedagogy and other aspects of graduate school.

Qualifying Examination

The qualifying examination must be completed no later than six semesters after matriculation. In most graduate curricula in the department, this consists of a research proposal—often in the form of a grant application—which the student submits to their committee and subsequently defends in an oral presentation. In the  Cell & Molecular Biology and Ecology, Behavior, Evolution & Marine Biology curricula , this is preceded by a comprehensive written examination testing the student’s general background from coursework.

Dissertation & Final Oral Examination

Candidates shall demonstrate their abilities for independent study in a dissertation representing original research or creative scholarship. A prospectus for the dissertation must be completed and approved by the readers, the Director of Graduate Studies, and the Biology Department Chair. Candidates must undergo a final oral examination in which they defend their dissertation as a valuable contribution to knowledge in their field and demonstrate a mastery of their field of specialization in relation to their dissertation. All portions of the dissertation and final oral examination must be completed as outlined in the  GRS General Requirements for the Doctor of Philosophy Degree . The results of the dissertation must be presented at a department colloquium.

Graduation Timeline

Forms and additional information about PhD graduation can be found on the GRS website .

9 – 12 months before proposed graduation date

• Dissertation Prospectus & approval form due to Graduate Program Specialist for departmental review and submission to GRS

Semester prior to your intended graduation cycle

• Intent to Graduate Form completed online

About 2 months before dissertation defense

• Meet with Graduate Program Specialist, Director of Graduate Studies, and Faculty Advisor to review requirements for defense
• Arrange for Special Service Appointments if committee members are not BU faculty
• Send first draft of dissertation to readers

Once defense date is confirmed with committee

• Reserve room(s) for public seminar and defense

At least three weeks prior to dissertation defense

• Schedule of Final Oral Exam with Abstract Approval due to GRS
• Properly formatted draft of dissertation submitted as PDF to [email protected]

At least two weeks prior to dissertation defense

• Send dissertation to all committee members

At least one week prior to dissertation defense

• Send program information to Graduate Program Specialist

See the Graduate Program Guide for final dates to submit dissertation to ETD

• Submit final dissertation to ETD (online submission)
• Complete electronic signature page and forward confirmation to Graduate Program Specialist along with a copy of your submitted dissertation

MS Degree (En Route to PhD)

Option one: A PhD student who has advanced to candidacy (as demonstrated by passing the PhD qualifying exam), and has completed 32 credits of graduate-level coursework (not including research) may apply to the Graduate School for an MS degree in Biology. This must be approved by the Director of Graduate Studies within the Biology Department. The student’s major professor will receive notification of this application process.

Option two: A PhD student who has taken, but has not advanced to candidacy based on the PhD qualifying examination, may still receive an MS degree. This student may receive a Coursework MS degree provided they have completed 32 credits of coursework (not including research credits). Alternatively, this student may receive a Scholarly Paper or Research Thesis MS degree if the written portion of the qualifying examination is adapted to ensure it is of sufficiently high quality for a MS degree, and approved by a majority of the qualifying exam committee and the Director of Graduate Studies.

Interdisciplinary Study Options

Biology PhD students have the option to participate in the Boston University Graduate Program in Urban Biogeoscience and Environmental Health (BU URBAN), the National Science Foundation Research Traineeship Program Understanding the Brain: Neurophotonics (NSF NRT UtB: Neurophotonics), and the Biogeoscience Advanced Graduate Certificate Program . These programs require separate applications in addition to the standard Biology PhD application; those interested in BU URBAN are encouraged pre-apply .

Time Limits

Officially, the PhD must be completed within seven years after the first registration for doctoral study. PhD degrees are conferred in either May, August, or January, as specified on the GRS website . In addition, the PhD candidacy expires after the fifth anniversary of passing the Qualifying Examination. Petitions to extend this deadline are possible at the discretion of the Director of Graduate Studies, the Department Chair, and the Dean of the Graduate School, and can be obtained from the Office of the Graduate School of Arts & Sciences.

The Biology Department guarantees support for five years for all PhD students, contingent on satisfactory performance in the program. PhD students are encouraged to apply for fellowships and grants at funding agencies. All domestic students should apply for NSF Graduate Research Fellowships in the Fall semester of their first or second year. 

Travel Grants may be available to assist students in their travel to professional scientific meetings; students presenting papers or posters on their research will receive first consideration.

Common Types of Funding:

Dean’s Fellowships: These are non-service fellowships allocated to first-year PhD students that do not have immediate teaching requirements.

Teaching Fellowships: These provide a stipend plus full tuition and fees for up to four full courses per semester plus a 2-credit teaching course. Teaching responsibilities usually require approximately 20 hours per week. Full or partial awards may be given.

Doctoral Research Fellowships: These awards are given to students who assist individual faculty with specific areas of research. These Research Fellowships provide a stipend and full tuition. The supervising faculty member determines the specific duties of the Research Fellow.

In addition to the above funding sources, several competitive Department awards and fellowships are available to graduate students in the Department of Biology.

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PhD in Biology

The PhD program in Biology is a research-intensive program that also has a strong focus on teaching, designed to produce top scientists and educators with a broad base of knowledge to tackle the most important biological problems of today. This is accomplished through research training, graduate-level courses, seminars, and teaching experience. Entering students are assigned an advisory committee of three faculty members who work with the student to plan a suitable program based on the student's experience and interests.

PhD students in Biology receive full tuition support for six years, a competitive stipend, health coverage, and receive extensive training in pedagogy, outreach, and communication that prepares them for careers in academia, biotechnology, education, and policy. Students are required to serve as teaching assistants for at least two semesters. Most students complete the requirement for teaching experience in the first year by assisting in the introductory undergraduate biology courses for two semesters.

During the first year, students become familiar with research opportunities in the department by doing short research rotations with various faculty members. This experience in expected to lead to the selection of a research focus for the PhD thesis, and it also sets the stage for cross-disciplinary approaches to the thesis topic that is pursued.

First- and second-year students usually take some graduate courses that are selected in consultation with the advisory committee, in addition to one required course: either Biology 243: Topics in Molecular and Cell Biology; Biology 244: Topics in Evolutionary Ecology; or Biology 246: Topics in Physiology and Animal Behavior.

The department faculty are educators as well as researchers, and the graduate students benefit from this expertise. Graduate students serving as teaching assistants have the opportunity to take a course on teaching and pedagogy to improve their skills, or to become involved with upper level courses in their area of expertise. In addition, we offer a one-month summer program for PhD students interested in contributing to course design and honing their teaching skills via GIFT, the Graduate Institute For Teaching .

Research Concentrations

Browse the six concentration areas below for recommended programs of study and relevant courses:

• Biology Education Research
• Ecology, Behavior and Evolution
• Global Change Biology
• Genetics and Molecular Biology
• Developmental Biology
• Physiology, Neurobiology and Animal Behavior

Graduate credit for a course requires a grade of B- (B minus) or better. A list of biology department undergraduate and graduate courses can be found by selecting the Courses button from the top bar.

Throughout the year, graduate students benefit from a variety of seminars on current research that are presented by faculty, fellow graduate students, and invited speakers.

Organismic and Evolutionary Biology

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Organismic and Evolutionary Biology is one of the programs in the Harvard Integrated Life Sciences , which facilitates collaboration and cross-disciplinary research. Visit HILS for additional application instructions.

Research in organismic and evolutionary biology (OEB) involves studying biological processes that span a continuum from single cells to entire ecosystems, conducting field and laboratory studies that are key to understanding the evolution of organisms, how biodiversity is generated and maintained, how organisms work, and how organisms interact with their environment. 

You will have access to an impressive array of resources, from modern research facilities to extensive biodiversity collections in the Herbaria and Museum of Comparative Zoology, with the ability to conduct plant and animal research at the Concord Field Station, the Harvard Forest, and the Arnold Arboretum. Many students conduct extensive fieldwork around the world at locations as varied as Antarctica, Kenya, and Brazil.

As part of the program, you will benefit from OEB’s broad connections in the life and physical sciences, including the Broad Institute; the Departments of Molecular and Cellular Biology, Human Evolutionary Biology, Systems, Synthetic, and Quantitative Biology, Psychology, Earth and Planetary Sciences, and Physics; and the Center for Brain Science and Broad Institute, as well as the Harvard John A. Paulson School of Engineering and Applied Sciences.

Examples of student dissertation topics include “The Structure, Function, and Evolution of the Fern Vascular System,” “Adaptations to Life on an Oxidizing Planet: Insights from the Evolutionary Ecophysiology of Iron-Respiring Bacteria,” and “High-Throughput Functional Characterization of Regulatory Variants Related to Human Evolution and Disease.”

Graduates have secured academic positions at institutions like Yale University, Princeton University, Stanford University, and the University of Chicago. Others have begun their careers with leading organizations like Ginkgo Bioworks, Whole Biome, and USDA Health.

Additional information on the graduate program is available from the Department of Organismic and Evolution Biology and requirements for the degree are detailed in Policies .

Admissions Requirements

Please review admissions requirements and other information before applying. You can find degree program-specific admissions requirements below and access additional guidance on applying from the Department of Organismic and Evolution Biology.

Standardized Tests

GRE General: Not Accepted 

Contacting Faculty

Students do not normally do rotations, but instead are admitted to work with a specific faculty member. Before applying, prospective students should review the department’s research page and reach out directly to their faculty of interest with a personal note, including a CV and a description of their research experiences and interests. Faculty may ask to speak on the phone or by Zoom or may extend an invitation to visit campus before the admissions cycle begins. These contacts are critical as part of the application process.

Theses & Dissertations

Theses & Dissertations for Organismic and Evolutionary Biology

See list of Organismic and Evolutionary Biology faculty

APPLICATION DEADLINE

Questions about the program.

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School of Biological Sciences

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Ph.d program in biology.

Our PhD program accepts students possessing Bachelors or Masters degrees who are planning a career in science. The purpose of our PhD program is to prepare highly qualified scientists who have excellent up-to-date training in the fields of biology or bioinformatics, who are actively involved in scientific research, are capable of making significant contributions to their scientific field, possess all the necessary skills for effective oral and written communication with colleagues, and can successfully compete in the job market for postdoctoral and research scientist level positions in academia and science-related industries. We currently have approximately 100 PhD students in the Biology PhD program of the School of Biological Sciences. Of recent PhD graduates, 70% are currently employed as postdoctoral researchers in academia, 9% as government scientists, 9% as industry scientists, 4% as instructors, and 9% are enrolled in further professional training. The average number of journal publications is 3.9 per student and the average number of presentations at conferences around the world is 4.3 per student.

Areas of concentration:

• Ecology, Evolution, and Behavior   including population and evolutionary ecology; community ecology; aquatic chemical ecology; ecological genomics; sensory ecology; evolution of development, behavior, and sociality; biological oceanography; environmental microbiology; theoretical ecology.
• Molecular and Cell Biology   including eukaryotic and prokaryotic cell biology; molecular physiology; molecular biophysics and structural biology; animal, plant, and microbial molecular genetics; human genomics; molecular evolution.
• Computational Biology and Bioinformatics   (with available   M.S.   and interdepartmental   PhD   programs in Bioinformatics) focusing on DNA and protein sequence analysis; comparative genomic analysis; macromolecular structure modeling including protein-protein, protein-nucleic acid, and protein-small molecule interactions; molecular evolution.

Ph.D. Program Overview:

The PhD in Biology is a research-based degree involving deep immersion in a topic within biology with mentoring from an advisor and thesis committee with expertise in the field of study. Coursework is typically completed within the first 2 years (18 credit hours). Students who have previously earned an M.S. degree or taken graduate courses elsewhere can negotiate up to 9 transfer credit hours upon entering our program. Selection of a thesis advisor is made in discussion with our faculty and/or via lab rotations during the first year, and the composition of the thesis committee is established by the student by the end of the first year.

PhD students in the School of Biological Sciences are each supported by a stipend and do not pay tuition, only modest   fees   each semester. Stipend support comes from teaching or research assistantships which complement research training towards the PhD. Whether PhD stipends are earned from research or teaching assistantships after the first year is decided in consultation with a student's thesis advisor and the graduate committee. Our PhD stipend is $33,500. Several fellowships are available which can supplement the PhD stipend. 

Further Funding: Please take some time to review our   funding opportunities and fellowships .

Choosing between the Masters and Ph.D. Program: Our graduate program is primarily focused on the PhD which prepares students for careers in scientific research and employment in academia, industry, or government. We encourage potential applicants to the PhD program to directly contact individual   faculty   members whose research may appeal to them to discuss research interests and future opportunities. Students who are not yet sure of their interest in scientific research or are interested in other kinds of professional development should consider the M.S. degree in   Biology   or Bioinformatics . However, we accept only a small number of M.S. Biology students each year - typically those in special career circumstances with a prior relationship to our program and faculty. Admission decisions are made by our graduate committee (composed of Biology faculty) in consultation with all of the faculty in the School of Biological Sciences.

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Biology, PHD

On this page:, at a glance: program details.

• Location: Tempe campus
• Second Language Requirement: No

Program Description

Degree Awarded: PHD Biology

The PhD program in biology offers individualized courses of study tailored to students' interests that include laboratory, field and theoretical work. Flexibility in the program is achieved by requiring only one core class, which is a choice between two topics that cover the breadth of the research directions in this program.

Electives are also as flexible as possible and include areas outside of biology. The major goal is to provide students with the opportunity to create an adaptable and general degree program that allows both traditional and interdisciplinary approaches in any area of biology.

Program Faculty   PhD Students

• BIO 792 Research
• BIO 514 Statistical Models for Biology
• BIO 522 Populations: Evolutionary Ecology
• BIO 507 Advanced Evolutionary Medicine
• BIO 521 Landscape Ecology
• BIO 530 Scientific Teaching (required in first semester for students that will TA at any point in degree)
• BIO 533 K-12 STEM Education & Outreach
• BIO 539 Computing for Research
• BIO 541 SOLS Seminar Series
• BIO 542 SOLS Current Topics in the Life Sciences
• BIO 591 Society and Natural Resource Management
• BIO 591 Social-Ecological Systems & Adaptation
• BIO 591 Drylands in a Changing Earth
• BIO 591 Population Genetic Reading Group
• BIO 591 Communication for Scientists
• BIO 591 Genetics and Genomics of Behavior
• BIO 620 Research Prospectus Writing
• EVO 501 Current Topics in Evolutionary Biology
• EVO 601 Principles of Evolution
• EVO 610 Research Areas of Evolution

Application and admission information

How to apply.

Applications open September 1 for admission in Fall of the following year. The application deadline is December 1 . We accept applications for Fall semesters only. We cannot guarantee that applications received after the December 1 deadline will be considered for admission.

All applicants must apply by filling out ASU's Graduate Admissions application. All application materials must be submitted through the application or to Graduate Admissions directly. Please do not mail or email any documents to the School of Life Sciences. 

Required materials and information include the following:

• 1-2 page personal statement
• An up to date CV or resume
• The names of relevant SOLS faculty you have been in touch with who you might be interested in being supervised by
• Unofficial transcripts and English proficiency test scores (if applicable)
• The names and emails of at least 3 recommenders to write you letters of recommendation

Application review process and timeline

Following the December 1 deadline, faculty will begin reviewing applications. Applicants should monitor their My ASU priority tasks to ensure there are no missing materials in their application.

Faculty will decide which applicants they would like to invite to our Graduate Recruitment Weekends (GRWs), typically held in February. Applicants will hear from the School of Life Sciences in January if they are invited to participate in the GRWs.

Admission decisions will begin after the GRWs, and applicants typically receive final decisions by April 1.

Requirements

Minimum requirements for admission include the following:

• Cumulative GPA of at least 3.0 on a 4.0 scale
• There are other ways to demonstrate English proficiency beyond the tests, so please refer to ASU's English proficiency webpage to review how you might satisfy requirements

Desired qualifications typically seen in competitive candidates:

• Research experience and a letter of recommendation from a faculty research supervisor
• English proficiency scores that meet these teaching assistant language proficiency requirements

Please note that the GRE is not required.

Students offered admission to a PhD program in the School of Life Sciences will typically receive a funding offer as well. While individual funding offers may differ to some degree, they typically include teaching assistant and/or research assistant positions each semester (summer optional) for 5 years. These positions provide financial coverage through the following:

• A standard salary stipend paid biweekly
• Tuition remission covering enrollment in 6-18 credit hours for fall and spring semesters and 1-14 credit hours for summer semesters
• Health insurance coverage

To discover more, check out the ASU Graduate College's funding opportunities !

Degree Requirements

84 credit hours, a written comprehensive exam, an oral comprehensive exam, a prospectus and a dissertation

Required Core (3 or 4 credit hours) BIO 514 Statistical Models for Biology (4) or BIO 620 Research Prospectus Writing (3)

Electives (67 or 68 credit hours)

Other Requirements (1 credit hour) BIO 610 Introduction to Responsible Conduct of Research in Life Sciences (1)

Culminating Experience (12 credit hours) BIO 799 Dissertation (12)

Additional Curriculum Information The program is tailored to the needs of the individual student. The plan of study is developed by the student and a supervisory committee consisting of a major professor and three additional faculty members. The electives credit hours are comprised of coursework showing breadth in biology subdisciplines, seminars and research.

An oral comprehensive examination is required in order to advance to candidacy, and a formal defense of the final written dissertation is required prior to graduation.

Admission Requirements

Applicants must fulfill the requirements of both the Graduate College and The College of Liberal Arts and Sciences.

Applicants are eligible to apply to the program if they have earned a bachelor's or master's degree in biology or a related discipline from a regionally accredited institution.

Applicants must have a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in the last 60 hours of their first bachelor's degree program, or a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in an applicable master's degree program.

Applicants must submit the following:

• graduate admission application and application fee
• official transcripts
• academic record form
• personal statement
• curriculum vitae or resume
• three letters of recommendation
• proof of English proficiency

Additional Application Information An applicant whose native language is not English must provide proof of English proficiency regardless of their current residency.

Research experience is desired for entry into this program.

Next Steps to attend ASU

Learn about our programs, apply to a program, visit our campus, learning outcomes.

• Able to communicate the findings and originality of their research, both orally and in writing.
• Able to perform research that they have designed to address important and challenging questions in the field of biology.
• Able to review scholarly literature as they develop specialized expertise in their selected research area.

Career Opportunities

Those who have earned a doctorate in biology are prepared for academic careers at every level, from community colleges to research universities. Their skills and knowledge are also valuable for government careers with federal and state agencies responsible for wildlife management and conservation, and for careers in industry and nongovernmental organizations.

Career examples include:

• food, agriculture and health care scientists in academic, private and industrial labs
• principal investigators in government labs and nonprofit organizations
• professors or instructors in universities and colleges
• science teachers in elementary and high schools
• wildlife, animal and conservation scientists

Program Contact Information

If you have questions related to admission, please click here to request information and an admission specialist will reach out to you directly. For questions regarding faculty or courses, please use the contact information below.

• [email protected]
• 480/965-1768

Department of Biology

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• Ph.D. Program

The Department of Biology offers a Doctor of Philosophy degree in Quantitative Biology with research emphasis in a number of areas including: Genome Biology & Genetics, Cell Biology, Ecology & Evolutionary Biology, or Microbiology. The doctoral program is designed to train students to apply sophisticated quantitative techniques to solving basic and applied problems in biology. Students in this program will attain substantially greater quantitative skills than in traditional doctoral programs in the biological sciences, providing them with a competitive advantage in business, industry, government, and academia.

The  University Catalog  contains all UTA admission requirements for the Department of Biology Ph.D. program.

How to Apply

First, read the general instructions for applying to the graduate program and download the checklists available on the Office of Graduate Studies webpage:  www.uta.edu/admissions/graduate/apply

International applicants have specific requirements, and a separate admissions checklist, available here: https://www.uta.edu/admissions/apply/international-graduate  

Additionally, the Department requests that applicants fill out the following questionnaire as part of the application package:  https://common.forms.uta.edu/view.php?id=2034157   Students are accepted into our Ph.D. program as either direct admit or rotation students. 

Direct admit: To qualify as a direct admit, an applicant must secure the backing of a faculty member before applying, mention their faculty support in their letter of intent, and ask that their faculty member reach out to Dr. Mark Pellegrino ( [email protected] ).

To secure the support of a faculty member before applying, begin by visiting our Biology faculty page: https://www.uta.edu/academics/schools-colleges/science/departments/biology/faculty - research/faculty , review the research focuses of the labs, and contact individual faculty members to discuss your research interests. Explain how your interests align with their lab and inquire if they would consider having you join their lab as a new Ph.D. candidate.

Rotation program: Applicants would choose this option if they are unsure which area of research interests them. Please indicate your interest in the rotation program in your letter of intent. Applicants admitted into the program will perform three rotations in three separate labs. At the end of the rotation period, applicants are placed in a home lab under the guidance of a Rotation Program Committee. Applicants will select their rotation labs only after being admitted into the Ph.D. program. Please note that applications to the rotation program are only considered for the Fall semester . For more information about the rotation program, please contact the chair of the Rotation Program Committee, Dr. Mark Pellegrino ( [email protected] ).

ALL APPLICATIONS ARE REVIEWED ON THE FOLLOWING SCHEDULE:

Questions? Please contact : [email protected]  

ADMISSION REQUIREMENTS

Students interested in pursuing a Ph.D. in the Department of Biology must apply to either the “B.S. to Ph.D. track” or the “Ph.D. program”, with the corresponding prerequisites:

B.S. to Ph.D. track:  A bachelor's degree in biology, or a bachelor's degree in some other discipline with at least 12 hours of advanced level coursework (junior or senior level courses) in biology. This is the Ph.D. track designed for students with a B.S and without master’s degree, which tends to be the majority of applicants.

Ph.D. program:  This track is for students that already have a master's degree in biology, or at least 30 hours of graduate level coursework in biology. A minimum undergraduate GPA of 3.0 (4.0 scale), as calculated by the Graduate School, is expected. An applicant's GPA in science courses will be taken into consideration. If an applicant has a master's degree, the GPA from their master's degree will also be considered. For applicants with up to 30 hours of graduate coursework but no master's degree, the GPA from those hours will be also be taken into consideration.

GRE:   The Graduate Record Exam (GRE) is no longer required for admission. 

Application, scores, and transcripts:   Your online application will include instructions for uploading transcripts, test scores, and any other required materials. 

International students:  Students whose native language is not English must provide a score on the Test of Spoken English (TSE) of at least 45, a minimum score of 23 on the Speaking portion of the TOEFL iBT exam or a minimum score of 7 on the Speaking portion of the IELTS exam. See the following website for complete details:  https://www.uta.edu/admissions/apply/international-graduate

Letters of recommendation:  Three letters  of recommendation are required for admission to the program. Ideally one or two letters should come from professors or instructors who can speak to your academic abilities; others may come from employers or other individuals who can comment on your work ethic and potential for graduate study. Letters should be emailed directly from the letter writer to [email protected] . If you have questions regarding letters of recommendation, please email [email protected] and include your UTA ID #, provided within your online application.

Letter of intent:  Please email a Letter of Intent to [email protected] and include the following information:

• Your full name and UTA ID #.
• Indicate that you are seeking the Ph.D. degree in Quantitative Biology
• Tell us which research track or types of research you are interested in. Examples include: Genome Biology & Genetics, Cell & Developmental Biology, Ecology & Evolutionary Biology, or Microbiology
• Provide names of any specific faculty member(s) that you are interested in working with if you have particular preferences. Note, however, that it is also OK if you are not sure what faculty member you are interested in working with (which is common), in which case, we will use your research interests above to help match you with potential faculty research mentors. We also encourage you to reach out to faculty members directly by email to discuss the potential for research in their lab, and if you have contacted any, please put this information in your letter of intent.
• How do you decide on which lab to conduct research in for your PhD? - Some students have a clear idea of their research interests, others may not. Both situations are common and our PhD program can account for both types of students, and students can decide on a single lab upfront to conduct research (direct admission) or decide to rotate in multiple labs during their first year, and decide which of these they will do their PhD work in in the second year. Our diverse department enables a broad range of research opportunities.
• Identify if you wish to be admitted directly to a single faculty member’s laboratory (and identify this faculty member), or indicate that you are interested in potentially rotating in multiple labs (2-3 labs) in the first year to make this decision. If interested in a lab rotation, please also indicate which faculty members’ labs are of interest, and if you have been in contact these faculty. More information about Rotations is provided below.
• Your Letter of Intent should also include details of your experience, training, any research experience you may have had, and motivations that you believe make you a strong candidate for success in a PhD program. You may also briefly discuss the long-term professional goals that motivate you to work towards a PhD. This is also an appropriate place to clarify details of your academic and work experience to date.

What is the letter of intent used for:

• The letter of intent is used to help the department link interested candidates with potential research professors/labs, and serves as an opportunity for the applicant to make a personal argument for why they believe they will be successful in conducting a PhD based on their experience and motivations. Your letter will be forwarded to the faculty members you indicate, but you are encouraged to also contact those faculty members directly (e.g., by email) to determine if they are accepting new students. It is therefore useful to also visit the listed  faculty pages  to find faculty research that you might be interested in and contact those faculty members.

PhD Funding:  Ph.D. students are guaranteed 5 years of financial support in the form of Graduate Teaching Assistantships (GTAs), funded by the department, or Graduate Research Assistantships (GRAs), funded by grants to individual faculty members. Many students are supported by a mixture of these two assistantships throughout their PhD. All Ph.D. students enrolled full-time and receiving an assistantship also receive reduced-cost in-state tuition regardless of residence, and a major portion of these tuition costs are supported by the assistantships.

Checking the status of your application:   You can check your application and admissions status through the MyMav student portal, where you can also review if you have any missing items needed to complete your application.   CHECK YOUR STATUS IN MYMAV . To confirm receipt of letters of recommendation, first check with your recommenders before contacting  [email protected] .  Please include your UTA ID #, available via MyMav, in any correspondence with the Biology Department.

Lab Rotations

The UTA Biology Department encompasses numerous laboratories that conduct research on a remarkably broad range of topics, and choosing a lab and research topic can be difficult. About half of entering students join a single lab as a direct admission. The other portion of students decide to rotate in multiple labs their first year. If several labs are of interest to you, consider signing up for lab rotations your first semester. Our lab rotation option allows you to experience multiple labs before making a final decision. However, please be sure to reach out to at least two potential rotation faculty to be sure they are able and interested in taking students for rotation. In your letter of intent (see  How To Apply  above), inform the Biology Department which labs are of interest to you (list up to three labs), and note which faculty have agreed to host you as a lab rotation student if you are accepted.

• If you decide to enter the PhD program as a rotation student, you can do ½ semester rotations in two host labs to get a sense of the research being conducted prior to making a final decision on which lab you will conduct your PhD research in.  
• For those students opting for a lab rotation, your letter of intent will be forwarded to the faculty members you indicate, but you should also contact those individuals directly to determine if they are taking new students and have the resources to support your research. During your first semester you will join/rotate through up to three labs in which you are interested and that are currently recruiting students. If no fit is found during your first semester, you have the option of rotating through additional labs in your second semester. Please visit the listed faculty pages to find faculty research that you might be interested in and contact those faculty members to set up a rotation. We feel confident that you will find an exciting and dynamic dissertation lab to join.  

BIOLOGY GRADUATE HANDBOOK [PDF]

The Biology Graduate Handbook is currently under revision. Up-to-date information regarding the PhD in Biology is available in the UTA Graduate Catalog . Questions regarding the Biology graduate program should be emailed to [email protected] .

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Location : Life Sciences Building, Room 337, 501 S. Nedderman Dr., Arlington, TX 76019 Mailing address : P.O. Box 19498 Phone : 817-272-2872

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Human Genetics and Genomics, PhD

School of medicine, ph.d. program.

The Johns Hopkins Human Genetics Training Program provides a training in all aspects of human genetics and genomics relevant to human biology, health and disease. 

Advances in human genetics and genomics continue at an astounding rate and increasingly they are being integrated into medical practice. The Human Genetics Program aims to educate highly motivated and capable students with the knowledge and experimental tools that will enable them to answer important questions at the interface between genetics and medicine. Ultimately, our trainees will be the leaders in delivering the promise of genetics to human health.

The overall objective of the Human Genetics program is to provide our students with a strong foundation in basic science by exposure to a rigorous graduate education in genetics, genomics, molecular biology, cell biology, biochemistry and biostatistics as well as a core of medically-related courses selected to provide knowledge of human biology in health and disease. 

This program is also offered as training for medical students in the combined M.D./Ph.D. program.  Students apply to the combined program at the time of application to the M.D. program. (See section entitled Medical Scientist Training Program).

Research Facilities

Research laboratories are well equipped to carry out sophisticated research in all areas of genetics. The proximity to renown clinical facilities of the Johns Hopkins Hospital, including the Department of Genetic Medicine, and Oncology Center provides faculty and students with access to a wealth of material for study. Computer and library facilities are excellent. Laboratories involved in the Human Genetics Program span Johns Hopkins University; consequently supporting facilities are extensive.

Financial Aid

The program is supported by a training grant from the National Institute of General Medical Sciences. These fellowships, which are restricted to United States citizens and permanent United States residents, cover tuition, health care insurance and a stipend during year one.  Once a student has joined a thesis lab, all financial responsibilities belong to the mentor.   Students are encouraged, however, to apply for fellowships from outside sources (e.g., the National Science Foundation, Fulbright Scholars Program, Howard Hughes Medical Institute) before entering the program.

Applicants for admission should show a strong academic foundation with coursework in biology, chemistry and quantitative analysis.   Applicants are encouraged to have exposure to lab research or to data science.  A bachelor's degree from a qualified college or university will be required for matriculation.  GREs are no longer required.

The Human Genetics site has up-to-date information on “ How to Apply .” For questions not addressed on these pages, please access the contact imformation listed on the program page: Human Genetics and Genomics Training Program | Johns Hopkins Department of Genetic Medicine (hopkinsmedicine.org) .

Program Requirements

The program includes the following required core courses: Advanced Topics in Human Genetics, Evolving Concept of the Gene, Molecular Biology and Genomics, Cell Structure and Dynamics, Computational Bootcamp,  Pathways and Regulation, Genomic Technologies, Rigor and Reproducibility in Research, and Systems, Genes and Mechanisms of Disease. Numerous elective courses are available and are listed under sponsoring departments.

Our trainees must take a minimum of four electives, one of which must provide computational/statistical training.

The HG program requires the “OPTIONS” Career Curriculum offered by the Professional Development and Career Office.  OPTIONS is designed to provide trainees with the skills for career building and the opportunity for career exploration as well as professional development training

Human Genetics trainees also take a two-week course in July at the Jackson Labs in Bar Harbor, Maine entitled "Human and Mammalian Genetics and Genomics: The McKusick Short Course" which covers the waterfront from basic principles to the latest developments in mammalian genetics. The faculty numbers about 50 and consists roughly in thirds of JAX faculty, Hopkins faculty and “guest” faculty comprising outstanding mammalian geneticists from other US universities and around the world.

The courses offered by the faculty of the program are listed below. All courses are open to graduate students from any university program as well as selected undergraduates with permission of the course director.

Trainees must complete three research rotations before deciding on their thesis lab.  They must also participate in the Responsible Conduct of Research sessions offered by the Biomedical Program; starting at year 3, students must attend at least two Research Integrity Colloquium lectures per year. 

Our trainees participate in weekly journal clubs, department seminars, monthly Science & Pizza presentations as well as workshops given twice a year on diversity, identity and culture.

At the end of the second year, trainees take their Doctoral Board Oral Examination.  Annual thesis committee meetings must be held following successful completion of this exam.

Average time for completion is 5.3 years.

Graduates from the Human Genetics program pursue careers in academia, medicine, industry, teaching, government, law, as well the private sector.  Our trainees are encouraged to explore the full spectrum of professional venues in which their training my provide a strong foundation. Driven by curiosity and a desire for excellence, our trainees stand out as leaders in the chosen arenas of professional life. They are supported in the development of their career plans by a program faculty and administration who are dedicated to their success, and by a myriad of support networks across the Johns Hopkins University, many of which are provided by the Professional Development Career Office of the School of Medicine.

Best Global Universities for Biology and Biochemistry in the United States

These are the top universities in the United States for biology and biochemistry, based on their reputation and research in the field. Read the methodology »

To unlock more data and access tools to help you get into your dream school, sign up for the  U.S. News College Compass !

Here are the best global universities for biology and biochemistry in the United States

Harvard university, massachusetts institute of technology (mit), stanford university, university of california berkeley, university of california san francisco, university of california san diego, university of washington seattle, johns hopkins university, university of california los angeles, cornell university.

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• # 1 in Best Universities for Biology and Biochemistry
• # 1 in Best Global Universities

Founded in 1636, Harvard University is the oldest higher education institution in the U.S. The bulk of Harvard's... Read More

• # 2 in Best Universities for Biology and Biochemistry
• # 2 in Best Global Universities

Massachusetts Institute of Technology, founded in 1861, is located in Cambridge, Massachusetts, near Boston. Around... Read More

• # 3 in Best Universities for Biology and Biochemistry
• # 3 in Best Global Universities

Stanford University was founded in 1885 and is located in California’s Bay Area, around 30 miles south of San Francisco... Read More

• # 4 in Best Universities for Biology and Biochemistry
• # 4 in Best Global Universities

The University of California—Berkeley is situated roughly 15 miles from San Francisco in what is known as the Bay Area... Read More

• # 5 in Best Universities for Biology and Biochemistry
• # 16 in Best Global Universities  (tie)

The University of California—San Francisco is a public institution that was founded in 1864. The health sciences-focused... Read More

• # 7 in Best Universities for Biology and Biochemistry
• # 20 in Best Global Universities

The University of California—San Diego is a public institution that was established in 1960. It is located in the La... Read More

• # 8 in Best Universities for Biology and Biochemistry
• # 6 in Best Global Universities

The University of Washington is a public institution that was founded in 1861. The school's oldest and largest campus in... Read More

• # 10 in Best Universities for Biology and Biochemistry
• # 10 in Best Global Universities

Johns Hopkins University is a private institution that was founded in 1876. The school has campuses located in and... Read More

• # 12 in Best Universities for Biology and Biochemistry
• # 14 in Best Global Universities

The University of California—Los Angeles, commonly referred to as UCLA, is a public institution that was founded in... Read More

• # 13 in Best Universities for Biology and Biochemistry
• # 21 in Best Global Universities

Cornell University is a private institution that was founded in 1865. The Ivy League school is located in Ithaca, New... Read More

Ph.D. Programs

Phd programs in the department of biology.

The Department of Biology has PhD Degrees in the following areas: Cell and Molecular Biology, Ecology and Evolutionary Biology.

You can find specific information about each program in the links below

PhD in Cell and Molecular Biology

PhD in Ecology

PhD in Evolutionary Biology

You can find specific application information about each program in the links below

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Diversity, Equity, and Inclusion

Message from the director.

A welcome message from Sabrina Diano, PhD, Director, Institute of Human Nutrition.

IHN alumni are advancing nutrition around the world through work in the health care industry, clinical research, medical education, and more.

Research Laboratories

Learn more about the Institute for Human Nutrition's research.

In the Community

Local initiatives.

IHN is a proud supporter of local organizations and activities here in Washington Heights and the surrounding communities.

PhD in Nutritional and Metabolic Biology

The Nutritional and Metabolic Biology (NMB) PhD training program prepares students to work at the frontiers of biomedical research in nutritional and metabolic sciences, exploring the role of nutrition in maintaining optimal human health.  The objective of the training program is to prepare individuals who will conduct original basic science research, teach in medical schools and universities, and hold positions of leadership in community and international nutrition.

Housed within the Institute of Human Nutrition (IHN) at Columbia University Medical Center (CUMC), this inter-disciplinary and multi-departmental training program is highly structured and comprises both coursework and basic research. The NMB program is one of the few pre-doctoral training programs in nutrition in the United States that is located within a medical school and is unique among the other PhD programs at CUMC with an equal number of MDs and PhDs as faculty mentors (including ten MD/PhDs). The location of the NMB training program in a medical school offers trainees a wide array of research opportunities in laboratories headed by established senior scientists as well as NIH-funded younger independent investigators, all focused on the role of nutrition and metabolism in health and disease.

nmb_phd_program_faculty_-_life_at_columbia

nmb_phd_program_students_-_life_at_columbia

For information on NMB faculty, please visit the Faculty page on the Graduate School of Arts and Sciences (GSAS) site .

Lori Zeltser, PhD

• Co-director

Anthony Ferrante Jr., MD, PhD

College of Biological Sciences

Department of Neurobiology, Physiology and Behavior

The Department of Neurobiology, Physiology and Behavior is a vibrant academic unit that investigates the fundamental processes shared by all animals. These include growth, reproduction, movement, response to stimuli, and maintenance of homeostasis. Faculty explore the highly integrated and precisely regulated physiological mechanisms underlying these vital functions. The department’s eponymous major offers students an exciting blend of hands-on labs, foundational courses in biological sciences, chemistry, mathematics, and physics and advanced topics like animal behavior, physiology of specific organ systems, developmental neurobiology, and endocrinology. Graduates pursue diverse career paths, including roles in medical professions, research and academia, and clinical settings. 

Undergraduate Programs

Graduate programs, affiliated centers.

Center for Neuroscience

The Center for Neuroscience is dedicated to understanding all levels of brain development, function and dysfunction. These discoveries help provide the foundation for improvements in health and new therapies for disease. 

Center for Mind and Brain

The Center for Mind and Brain (CMB) is devoted to supporting the collaborative work of world-class scientists from a broad spectrum of disciplines encompassing divergent social, biological, computational and medical perspectives.

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Doctor of Philosophy (PhD)

Offered By: Department of Molecular Microbiology and Immunology

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About the PhD in Molecular Microbiology and Immunology Program

Through a departmental core curriculum as well as research area-specific courses, laboratory rotations, qualifying examinations, mentored research, and professional development, all MMI PhD students are prepared to engage in cutting edge research and scholarship that advances knowledge. MMI PhD students hone their scholarship, research and professional skills. Course and laboratory work can address problems in microbial pathogenesis, immunology, disease transmission, and diseases related to malaria, mosquito and arboviral biology.

Students can choose to complete the traditional MMI PhD program or the MMI PhD program concentration in Rigorous, Reproducible, and Responsible Research Investigation in Immunology & Microbiology (R 3 IM). The concentration in Rigorous, Reproducible, and Responsible Research Investigation in Immunology & Microbiology (R 3 IM) conveys a broad background in immunology and infectious diseases research, with a special emphasis on critical thinking, logic, ethics, and written and oral skills to help graduates become communicators of complex scientific concepts and agents of change in their workplaces and communities.

What Can You Do With a Graduate Degree In Molecular Microbiology And Immunology?

Sample careers.

• Postdoctoral Fellow, Research Associate
• Science Communicator or Writer
• Scientist (academia, industry)
• Specialist in Science Regulatory Affairs
• Faculty, Professor
• Science Advocate (nonprofit agencies)
• Public Health Service (NIH, CDC, FDA)
• Science Policy Fellow

Curriculum for the PhD in Molecular Microbiology and Immunology

Browse an overview of the requirements for this PhD program in the JHU  Academic Catalogue  and explore all course offerings in the Bloomberg School  Course Directory .

Current students can view the MMI student handbook on the MMI portal site.

Topic Areas

MMI faculty are recognized experts in a wide variety of infectious diseases research areas, allowing our PhD students to study the biology of disease based on their research interests and career goals. Our PhD students gain a comprehensive understanding of infectious diseases that provides a foundation to launch careers that directly tackle critical public health challenges.  

Topic Areas include: 

• Bacterial pathogenesis
• Cell biology 
• Fungal pathogenesis 
• Medical entomology/Disease ecology
• Parasite pathogenesis 
• Rigor, Reproducibility, and Responsibility in Scientific Practice
• Tick-borne diseases
• Vaccine development 
• Vector biology
• Viral pathogenesis

Admissions Requirements

For general admissions requirements, please visit the How to Apply page.

Standardized Test Scores

Standardized test scores (GRE) are  optional  for this program. The admissions committee will make no assumptions if a standardized test score is omitted from an application, but will require evidence of quantitative/analytical ability through other application components such as academic transcripts and/or supplemental questions.  Applications will be reviewed holistically based on all application components.

Vivien Thomas PhD Scholars

The  Vivien Thomas Scholars Initiative (VTSI)  is an endowed fellowship program at Johns Hopkins for PhD students in STEM fields. It provides full tuition, stipend, and benefits while also providing targeted mentoring, networking, community, and professional development opportunities. Students who have attended a historically Black college and university (HBCU) or other minority serving institution (MSI) for undergraduate study are eligible to apply. To be considered for the VTSI, you will need to submit a SOPHAS application, VTSI supplementary materials, and all supporting documents (letters, transcripts, and test scores) by December 1, 2023. VTSI applicants are eligible for an  application fee waiver , but the fee waiver must be requested by November 15, 2023 and prior to submission of the SOPHAS application.

All full-time PhD students will receive the following support for all years of the program: stipend, full tuition, individual health insurance, University Health Services clinic fee, vision insurance, and dental insurance.

Need-Based Relocation Grants Students who  are admitted to PhD programs at JHU  starting in Fall 2023 or beyond can apply to receive a $1500 need-based grant to offset the costs of relocating to be able to attend JHU.   These grants provide funding to a portion of incoming students who, without this money, may otherwise not be able to afford to relocate to JHU for their PhD program. This is not a merit-based grant. Applications will be evaluated solely based on financial need.  View more information about the need-based relocation grants for PhD students .

Questions about the program? We're happy to help.

Alex Kim , Senior Academic Program Coordinator  Department of Molecular Microbiology and Immunology Johns Hopkins Bloomberg School of Public Health 615 North Wolfe Street, Room E5014 Baltimore, Maryland 21205 Fax: (410) 955-0105

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Dr. Lisa Korn Receives 2024 Dostanic Award

Lisa Korn, MD, PhD , instructor of medicine (rheumatology, allergy and immunology) has been selected to receive the 2024 Iva Dostanic, MD, PhD, Physician-Scientist Trainee Award.

The award is named in honor of the late Iva Dostanic, MD, PhD, a trainee who passed away in 2011 from ovarian cancer, who is described as having been an extraordinarily outstanding physician-scientist.

Presented annually to a physician-scientist trainee in the Department of Internal Medicine, the award has been given to more than a dozen trainees, including Dostanic herself, since its inception in 2011.

“I feel enormously honored to receive this award in memory of Iva Dostanic, who was known as a brilliant scientist and an amazing person,” Korn said.

I feel enormously honored to receive this award in memory of Iva Dostanic, who was known as a brilliant scientist and an amazing person. Lisa Korn, MD, PhD

Korn was recruited to the department in 2015 for the ABIM Physician-Scientist Research Pathway. She completed a clinical fellowship in rheumatology in 2019 and worked as a postdoctoral fellow in the laboratory of Ruslan Medzhitov, PhD, Sterling Professor of Immunobiology. Most recently, she joined the section of Rheumatology, Allergy and Immunology as an instructor in 2022.

Medzhitov and Section Chief Richard Bucala, MD, PhD , Waldemar Von Zedtwitz Professor of Medicine (Rheumatology) and professor of pathology, Yale School of Medicine, and of epidemiology (microbial diseases), Yale School of Public Health, nominated Korn for the award, citing her research and clinical acumen.

“She displays an impressive ability to reach outside her prior expertise to pursue the most interesting questions, and we believe that her discoveries will ultimately have a large impact on introducing a more rational, scientific evidence-based way to approach diet in the treatment of multiple disorders,” they wrote.

She displays an impressive ability to reach outside her prior expertise to pursue the most interesting questions, and we believe that her discoveries will ultimately have a large impact on introducing a more rational, scientific evidence-based way to approach diet in the treatment of multiple disorders. Richard Bucala, MD, PhD and Ruslan Medzhitov, PhD

Peter Aronson, MD , C.N.H Long Professor of Medicine (nephrology), has chaired the committee that selects the awardee since it was formed. During the selection process, he and the committee members noted Korn’s many achievements.

“The committee was impressed with the high quality of her research and publications on regulation of immune cells in the intestine during both her MD-PhD training and her postdoctoral training at Yale,” Aronson said. “In addition, she has demonstrated outstanding success in competing for multiple career development grants.”

Korn’s research focuses on the influence of dietary constituents on mucosal immunity, a subject she was introduced to while working in Medzhitov’s laboratory. Interested in the interaction between the immune system and the environment, she is intrigued by the connection between the gut and rheumatic diseases, from inflammatory arthritis to lupus, she said.

“The immune cells in the gut are on the front line with the environment,” Korn explained. “There is just a single layer of cells and some mucus that separates them from all the food and bacteria within the gut.”

Fred Gorelick, MD , Henry J. and Joan W. Binder Professor of Medicine (digestive diseases) and Cell Biology, who specializes in acute inflammation of the pancreas, recalls reviewing Korn’s K08 grant.

“At first, I was surprised a rheumatologist was submitting a research grant on the intestine, but the intestine is a very important immune sensing organ,” Gorelick said. “Those immune cells control local responses in the intestine, and they can, by mechanisms that aren't entirely understood, affect other tissues.”

With a plethora of mentors at Yale, Korn considers herself fortunate to have a supportive group behind her, including Medzhitov, a renowned creative thinker, who continues to impact Korn’s work. In addition, Korn is grateful for her mentorship committee–Joe Craft, Fred Gorelick, Martin Kriegel, Noah Palm, and Andrew Wang–and for the clinic-related work she has done with Cristina Brunet. The members of the Section of Rheumatology, Allergy and Immunology are wonderful sources of scientific and clinical wisdom, Korn said.

Korn credits her late mother, Eda T. Bloom, PhD, for her love of science. Bloom passed away suddenly in 2008, while Korn was studying for her MD-PhD at the University of Pennsylvania.

“My first scientific role model was my mom, one of the first women in her department to receive a PhD in immunology,” Korn said. “I followed in her footsteps, and I’d like to think she’d be proud.”

Korn will present her research and receive the Dostanic Award at Medical Grand Rounds on June 20, 2024, which will be attended by Dostanic’s parents.

The Department of Internal Medicine at Yale is among the nation's premier departments, bringing together an elite cadre of clinicians, investigators, educators, and staff in one of the world's top medical schools. To learn more, visit Internal Medicine.

Featured in this article

• Lisa Korn, MD, PhD Instructor of Medicine (Rheumatology, Allergy & Immunology)
• Peter S. Aronson, MD C. N. H. Long Professor of Medicine (Nephrology) and Professor of Cellular And Molecular Physiology
• Richard Bucala, MD, PhD Waldemar Von Zedtwitz Professor of Medicine (Rheumatology) and Professor of Pathology and of Epidemiology (Microbial Diseases); Chief, Rheumatology, Allergy, & Immunology; Rheumatologist in Chief, Rheumatology; Affiliated Faculty, Yale Institute for Global Health
• Ruslan Medzhitov, PhD Sterling Professor of Immunobiology; Investigator, Howard Hughes Medical Institute
• Fred Sanford Gorelick, MD Henry J. and Joan W. Binder Professor of Medicine (Digestive Diseases) and of Cell Biology; Director Investigated Gastroenterology NIH T32, Internal Medicine; Deputy Director, MD-PhD Program
• Joseph Craft, MD Paul B. Beeson Professor of Medicine (Rheumatology) and Professor of Immunobiology; Paul B. Beeson Professor of Medicine, Internal Medicine: Rheumatology; Program Director, Investigative Medicine, Internal Medicine: Rheumatology
• Martin Kriegel, MD, PhD Associate Professor Adjunct
• Noah Wolcott Palm, PhD Associate Professor of Immunobiology
• Andrew Wang, MD/PhD, AB Associate Professor of Internal Medicine (Rheumatology)
• Cristina Brunet, MD Associate Professor of Medicine (Rheumatology, Allergy & Immunology); Arthritis and Rheumatology Clinic Site Director, Interventional Immunology Center

Related Links

• Irene Chernova is the 2023 Dostanic Award Recipient
• Goldman-Israelow, Shung are 2022 Iva Dostanic Award Recipients

Medical Grand Rounds: "Renal Replacement Therapy: Past, Present and Future"

Graduate Education

Office of graduate and postdoctoral education, phd defense by chris johnson, april 18, 2024.

In partial fulfillment of the requirements for the degree of

Doctor of Philosophy in Biology

School of Biological Sciences

Chris Johnson

Will defend his dissertation

Investigating the Mechanisms Underlying Metamorphosis in the Chordate Ciona robusta

18, 4, 2024

Krone Engineered Biosystems Building (EBB) CHOA Room 1005

https://gatech.zoom.us/j/93986623573?pwd=eURpcHRrcXFwUENmcWNXRFBKZlpFUT09

  Thesis Advisor:

Alberto Stolfi, Ph.D.

Georgia Institute of Technology

Committee Members:

Annalise Paaby, Ph.D.

Brian Hammer, Ph.D.

William Ratcliff, Ph.D.

Pengpeng Bi, Ph.D.

Department of Genetics and Center for Molecular Medicine

University of Georgia

Tunicates are our closes living invertebrate relative, and as such, have given us tremendous insight on tunicate/vertebrate development and evolution. For this presentation, I discuss two projects that are tied together and contribute to the ongoing investigation underlying metamorphosis in tunicates.

The first chapter discusses our work on the transcriptional regulation governing the pattering of cells comprising the three sensory adhesive papillae of the Ciona robusta larva. Through the discovery and use of novel genetic markers and reporter constructs, we visualized and tracked the differentiation of various papilla cell types which range from collocytes responsible for secreting adhesive material, to ciliated neurons initiating metamorphosis and axial columnar cells potentially possessing chemosensory and contractile properties. Using CRISPR/Cas9, overexpression, and pharmacological perturbations, we identified the molecular mechanisms underlying the specification of these cell types and their contributions to metamorphic processes.

The second chapter describes the transcriptional regulation of adult Ciona muscle cell fusion. In vertebrates, all skeletal muscles form multinucleated myofibers through myoblast fusion. In tunicates like Ciona,  only adult muscles form multinucleated fibers through cell fusion, while larvae have mononucleated muscles instead. I show that this is achieved through combinatorial regulation that activates the expression of a key myoblast fusion factor only in adult muscles.

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34 Best Medical schools in Moscow, Russia

Updated: February 29, 2024

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Below is a list of best universities in Moscow ranked based on their research performance in Medicine. A graph of 922K citations received by 111K academic papers made by 34 universities in Moscow was used to calculate publications' ratings, which then were adjusted for release dates and added to final scores.

We don't distinguish between undergraduate and graduate programs nor do we adjust for current majors offered. You can find information about granted degrees on a university page but always double-check with the university website.

1. Moscow State University

For Medicine

2. Moscow Medical Academy

3. Russian National Research Medical University

4. RUDN University

5. National Research University Higher School of Economics

6. Moscow Institute of Physics and Technology

7. National Research Nuclear University MEPI

8. Bauman Moscow State Technical University

9. N.R.U. Moscow Power Engineering Institute

10. Russian Presidential Academy of National Economy and Public Administration

11. Moscow State Pedagogical University

12. National University of Science and Technology "MISIS"

13. Mendeleev University of Chemical Technology of Russia

14. Finance Academy under the Government of the Russian Federation

15. Plekhanov Russian University of Economics

16. Moscow Aviation Institute

17. Moscow State Institute of International Relations

18. Russian State Social University

19. Russian State Agricultural University

20. National Research University of Electronic Technology

21. Russian State University of Oil and Gas

22. new economic school.

23. Russian State University for the Humanities

24. Moscow State Technological University "Stankin"

25. State University of Management

26. Moscow State University of Railway Engineering

27. Moscow Polytech

28. Moscow Technical University - MIREA

29. Moscow State Linguistic University

30. Moscow State Mining University

31. Moscow International University

32. Pushkin State Russian Language Institute

33. Russian State Geological Prospecting University

34. moscow state technical university of civil aviation.

Universities for Medicine near Moscow

Medicine subfields in moscow.