phd in germany medicine

Ph.D. Program Medical Research - International Health

The Ph.D. program Medical Research - International Health combines health-related research in the widest sense and development cooperation. Special focus lies in educating researchers who in turn become multipliers in low- and middle-income countries (LMICs) and thus the next generation of health professionals. Not only acquiring knowledge for one's own interest, but importing and multiplying the knowledge to areas in need should be a guiding working motive while working as a Ph.D. student at CIH LMU . This motive is complying with global development strategies that have recognized the human resource crisis as a key factor in the deterioration of health systems in low- and middle-income countries.

From this stance arises a certain focus on candidates from LMICs to provide their home countries with competent and responsible researchers and health professionals. However, the Ph.D. program is open to all who share an interest in health-related issues in the field of development cooperation. Therefore, we encourage all professionals of any nationality to become part of the Ph.D. program, people working in health or health-related fields who are willing to improve the local research and health situation both through manpower and knowledge.

The Ph.D. program Medical Research - International Health is part of the Munich Medical Research School (MMRS), a central institution of the Ludwig-Maximilians-Universität München’s Medical Faculty. One of the main tasks of the MMRS is to establish Ph.D. core areas under the examination regulations for "Ph.D. Medical Research", in order to promote the faculty's internationalization. All doctoral students are automatically members of the MMRS upon admittance to the Ph.D. program Medical Research - International Health.

3 years full-time sandwich program

The doctoral project is supervised by both a local supervisor and two to three LMU supervisors. Doctoral candidates start their doctoral degree in Munich during Module Block I (Oct-Dec), but subsequently spend the research periods in their home country. After the first research period, Ph.D. candidates come to Munich for Module Block II (Jan-March) before completing the doctorate in their home country with the participation of the LMU supervisors

Doctor of Philosophy (Ph.D.) in Medical Research – International Health

Ludwig-Maximilians-Universität München, Germany

There are no tuition fees. However, please note that you need to be able to finance your research and the stays in Munich. Please see more details under "Costs" .

01 December 2023 to 31 January 2024, via the MMRS registration portal .

Our Ph.D. MR-IH study handbook contains important information about the course of study and the degree in a bundled format.

If you would like comprehensive information about the Ph.D. program or have specific questions, we offer an Information Session each year before the application period begins. The next information session will be announced in due time. Registration is required.

Further Information

• Research projects
• Supervision of your Ph.D.
• Ph.D. Thesis Submission
• Thesis evaluation and Oral Defense
• Obtaining the degree
• Scholarships

Application

Before applying, make sure to carefully read about the application requirements and selection process , and click here to register.

Helpful links for Ph.D. candidates

• Past Ph.D. Research Topics

The support by the German Federal Ministry of Economic Cooperation and Development is part of the EDCTP2 programme supported by the European Union.

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The LMU Medical Faculty has offered the Ph.D. Medical Research , a structured doctoral program of at least three years, for several years now. The Ph.D. is a research and training program, around medical research topics. In particular, it focuses on biomedical experimental, clinical, translational, and health sciences research. The Ph.D. Medical Research is designed for graduates who have completed an undergraduate degree (master's degree, state examination, or similar) in medicine, veterinary medicine, the natural sciences, pharmacy, a health science, or a related field and plan to conduct full-time research for at least three years .

In order to provide structured doctoral programs for Ph.D. students, university professors and lecturers in the Faculty of Medicine have joined together in various core areas and have committed themselves to adhere to the framework (recruitment, supervision, qualification and examination) specified in the study regulations for the Ph.D. Medical Research. This implies that events (workshops, seminars, etc.) are offered jointly within the framework of a core area. Depending on demand, doctoral students from other departments can also participate, which allows a lively exchange to take place and networks to be formed.

The concept of individual Ph.D. projects in the different research areas, coupled with a structured and partly integrative training program has proved successful over the past years.

Introduction of the Ph.D. Medical Research

A description of the Ph.D. Medical Research (structure, minimum requirements, etc.) can be found here.

Application and Selection

In order to obtain a Ph.D. in Medical Research, you must complete an application and selection process. Depending on which area you wish to carry out your Ph.D. in, there may be different application deadlines and procedures.

Core Areas and Research Training Groups

Currently, there are 5 Ph.D. core areas at the faculty, in addition to numerous SFBs and graduate programs that offer a Ph.D. in Medical Research. If none of the existing programs fit, there is the possibility of an "individual Ph.D. in Medical Research".

Supervision of your Ph.D. Project

Information on who can supervise your Ph.D. project, as well all documents required for this, can be found here.

Registration of your Ph.D. Project

Once you have successfully applied and been accepted for the Ph.D. Medical Research, you need to register yourself and your project using the online platform.

Curriculum and Courses

The Ph.D. Medical Research requires you to complete a curriculum comprising 30 ECTS. Information on how to accomplish this, can be found here.

Submission of your Ph.D. Thesis

Once you have completed all requirements for the Ph.D. Medical Research, you can submit your thesis.

Further Procedure and Oral Defence

Information on the evaluation process of your thesis, as well as on the oral defence can be found here.

Completion of the Procedure and Certificates

Once you have successfully completed your oral defence, you must submit the statutory copies to the University Library, before you can receive your degrees and certificates.

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PhD Programs

You are creative, curious and passionate about basic research? You would like to live in the oldest university town in Germany and work at one of the famous Max Planck Institutes? You would enjoy doing research in a supportive, stimulating and diverse environment?

We welcome graduate students from all over the world and a wide range of disciplines. Because Max Planck Institutes cannot confer PhD degrees, our students are simultaneously enrolled at Heidelberg University and receive their doctorate from one of the university’s faculties (  Biosciences Faculty ,  Chemistry Faculty ,  Physics Faculty ). These faculties have their own specific requirements, which students must fulfill.

PhD students at the institute perform research work and write a thesis under the supervision of a scientist who leads a research group in one of the institute’s departments. Students are usually enrolled in a graduate program. We offer access to a number of graduate schools, such as the  Max Planck School Matter to Life , the IMRPS-QD or other graduate schools of Heidelberg University. Most group leaders are associated with one or more of these graduate schools. Joining a graduate school makes you part of the Heidelberg life science community and gives you opportunities for scientific and social exchange outside of the institute.

For students who are not enrolled in a structured PhD program we offer support and guidance through individual Thesis Advisory Committees (TAC). The TAC monitors the progress and development of the PhD student and provides advice to both the student and his/her supervisor. The TAC members are also available for one-to-one consultations with the student and can become an important networking partner. Our PhD students can contact their committee members for personal discussions at any time.

For more information about the different PhD programs and the application procedure, please follow the links provided below. Alternatively, you may also directly contact the head of the department or group leader of the program you’re interested in. Just send an e-mail including your CV and a short description of your interests and expertise to establish first contact. Feel free to contact the PhD representatives or scientific coordinator with any general questions about working at our institute.

PhD Life at the Institute

Graduate Schools in Heidelberg

IMPRS Quantum Dynamics in Physics, Chemistry and Biology

Max Planck School Matter to Life

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PhD programme of the Faculty of Medicine

The Faculty of Medicine at Heinrich Heine University Düsseldorf offers the PhD in Medical Sciences. The structured doctoral programme is aimed at graduates of life science subjects.

The PhD in Medical Sciences covers important research areas of the Faculty of Medicine. These are assigned to three thematic clusters.

• Molecular Medicine for biomedical-experimental research projects
• Diagnosis and Therapy of Diseases includes the area of clinical and translational studies
• Health and Society includes projects in the health sciences

Accompanying advanced training events within the cluster curriculum deal with current research topics and enable a deepening of research methods with practical relevance. The training of doctoral students is rounded off by a basic curriculum with events on scientific core competences and key qualifications relevant to the profession.

Successful completion of the PhD programme leads to the award of the academic degree PhD in Medical Sciences or alternatively Dr. rer. med.

Apply with us and start your research career at the Faculty of Medicine of HHU!

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International phd program in medical life science and technology.

Welcome to the PhD program in Medical Life Science and Technology!

The PhD program “Medical Life Science and Technology” is a structured postgraduate program within the TUM School of Medicine aimed at students with a background in medicine, engineering or life sciences. The key research areas represented in the program are oncology , neuroscience , immunology/infection , biomedical imaging , cardiovascular research , and molecular medicine .

The main purpose of the program is to give students a thorough scientific training and the tools to become successful and independent scientists. Although the program is mainly focused on specialist knowledge, additional extra-disciplinary preparation complement the training. Skills gained from completing the PhD, including project management, analytical and communication skills, allow the PhD graduates not only to pursue a career in academia, but also to aim for a higher-level non-academic job in industry or healthcare.

The PhD program is part of the TUM Medical Graduate Center and the TUM Graduate School . Students enrolled in the PhD program become members of the Graduate School and benefit from its offers, such as financial support for international research activities .

Apply for our PhD program

Call opens: january 15, 2024 - application deadline: march 03, 2024, fully-funded phd positions in biomedical sciences.

Are you passionate about discovering the basis of health and disease, and applying your findings to answer clinically-relevant questions? Are you interested in one of the following topics?

Topic 1: Genes, Cells and Cell‑based Medicine Topic 2: Molecular Processes and Therapies Topic 3: Integrative Biomedicine

We are committed to meet today’s challenges in biology and medicine and realize that excellent science requires outstanding researchers. If you aspire to be one of them – we are looking forward to your application!

• An excellent scientific, international and collaborative learning environment 
• Individualised PhD supervision & mentoring
• Support of the thesis committee
• Advanced scientific, research-related & methods training
• Access to state-of-the-art infrastructure & core facilities
• Regular lectures, journal clubs & seminar series 
• Participation in one of our specialised research programs
• A structured program for personal & career development
• Participation in PhD symposia, retreats & social activities
• Travel grants for external courses & conferences
• Active alumni network

Students may, in addition, join our MDC-NYU PhD Exchange Program in Medical Systems Biology, iNAMES – MDC-Weizmann Helmholtz International Research School for Imaging from NAno to MESo, or become associated with the interdisciplinary HEIBRiDS Data Science School.

Who are we looking for?

To apply, a master’s degree (or equivalent) in in biology, biotechnology, medicine, molecular medicine, chemistry, computer science, data science, engineering, mathematics, physics or a related field , is required. You should hold or expect to obtain your degree not later than four months after the onsite interview week in Berlin (see below for the exact interview dates). Please check further details of your  Eligibility for application here .

Details on the  Recruitment process can be found here . Please make sure to check the FAQ section , as well.

Available projects

A list of participating group leaders and available projects, with detailed project descriptions, is available on our application portal upon registration.

How to apply? 

Calls for applications for central MDC recruitment are open twice a year, for Spring and Fall recruitment round . Applications are accepted exclusively via our online Application portal . Email applications for the central recruitment rounds can unfortunately not be accepted . Detailed information about the application process, including Application manual,  are available here . 

https://mdcphd.mdc-berlin.de

For the individual PhD vacancies potentially offered outside of the central Recruitment calls, please contact the scientific group leaders of your interest directly or follow the announcements in the  Open PhD positions section.  

For approximate timeline of the central Recruitment process, please see below. Please note that these are approximate dates, and they may vary from year to year. For the exact dates, follow the announcements on our page.

I have said […] that people go into science because they are misfits, and that it is a sheltered place where they can spin their own yarn and have recognition, be tolerated and happy, and have approval for it. Interview with Max Delbrück (1978), Oral History Project, California Institute of Technology Archives.

Spring round

• Call open: from November to January
• Online interviews: in March
• Starting date: latest 4 months from the onsite interview week (August same year)
• Call open: from May to July
• Online interviews: in September
• Starting date: latest 4 months from the onsite interview week (February next year)

PhD researchers are fully funded through an initial 3-year employment contract with the MDC. Salaries are competitive to national standards, according to 65% TVöD 13 ( monthly net payment of approximately 1800 EUR ). Health and social insurance are included in the contract. In many cases, contracts are extended for an additional year. All PhD researchers are expected to complete their doctoral research and thesis in 4 years. The university tuition fees per semester are approximately 350 EUR, and include public transportation ticket.

Int. PhD Program Brochure

https://www.mdc-berlin.de/system/files/document/PhD%20Booklet%202020%20FINAL_compressed_1.pdf

People in Focus Brochure

https://www.mdc-berlin.de/system/files/document/People%20in%20focus%20MDC%202020%20FINAL_compressed.pdf

Eligibility

Before starting your application, please ensure that you meet the eligibility requirements for the MDC International PhD Program. 

Please note that the eligibility criteria listed here apply exclusively to the MDC central Recruitment calls. Applications from candidates who do not meet the following requirements will not be considered for the central Recruitment calls (Fall and Spring Recruitment rounds).

Eligibility criteria for individual PhD vacancies offered outside of the central Recruitment calls may vary . For these, please contact the group leaders of your interest directly.

• Applicants are required to hold (or obtain within 4 months of the onsite interview week) a  university degree equivalent to a Master Degree (MSc)  in  biology, biotechnology, medicine, molecular medicine, chemistry, computer science, engineering, mathematics, physics, or some related field . If you have obtained your MSc degree outside of Germany, a German state authority, the Central Office for Foreign Education (ZAB), will need to confirm the recognition of foreign educational qualifications ("Statement of Comparability for Foreign Higher Education Qualifications”).
• Master degree equivalents include, for example, MD degree, Master of Research (MRes) and Master of Technology (MTech) degree, 4-yr BSc + 1-yr MSc programs, integrated programs such as Diploma or BSc-MSc integrated program, as well as BTech-MTech dual degrees. 
• Applicants holding only a BSc degree are not eligible to apply. Neither a BSc degree alone, even of a 4-year-long program, nor additional experience represent an equivalent to the MSc degree.
• Applicants still studying are welcome to apply, however, they should anticipate receiving their degree  not later than 4 months after the onsite interviews in Berlin .

Applicants are required to have some relevant research experience, gained through e.g. through bachelor and/or master research projects, lab rotations, study exchange programs, or in other relevant research projects,  either of experimental or theoretical nature.

To be eligible, a research master thesis should have been required for obtaining the MSc degree. There might be exceptions to this, e.g. for MD or pharmaceutical programs that do not require a final research thesis.  Research experience is, however, obligatory.

Required MSc final grade is ≥75%, or German score of at least 'good' (<2,5 of 5). To translate your final grades into German grading system, please use the  Modified Bavarian Formula Calculator . If your grades are lower than the required, the application will not be considered.

Only  early-stage researchers  are invited to apply: candidates should at the time of interviews be in the  first four years of their research careers  (i.e. master degree obtained within the last 4 years) and  should not have a doctoral degree already . If you have gained your master (or equivalent) degree more than four years prior to the interviews,  you are not eligible and your application will not be considered . 

Candidates who have  previously participated in the online interviews are not eligible to apply again.

Applicants need to have a  good command of written and spoken English , while German language skills are not required. English proficiency tests for non-native speakers are recommended (such as TOEFL test with a score of 95+ (internet-based), 240+ (computer-based) or 587+ (paper-based), IELTS certificate with a bandscore 7.0+, or Cambridge Certificate in Advanced English (CAE)), however,  the language proficiency certificates   are not a requirement for the application .

• Short-term interns (this includes BSc and MSc students) who have been working or registered at the MDC for  less than 3 months  during the 2 years prior to the online interviews are eligible to apply to the International PhD program recruitment,  if they apply to a different research group.
• Short-term interns ( less than 3 months ) who wish to  join the research group they have been affiliated before  (during the 2 years prior to the online interviews) are  not eligible for the International PhD Program   central   recruitment,  and are recommended to apply for  Internal PhD fellowship positions  (internal access only).
• MDC interns who have been working or registered at the MDC  for longer than 3 months  during the 2 years prior the online interviews,  are not eligible to apply to the International PhD Program recruitment.  Instead, they are considered  internal students  and may apply for  Internal PhD fellowship positions  (internal access only).
• If more than 2 years have passed between the end of affiliation (contract or guest registration) with the MDC and the online interviews, a candidate is eligible to apply for the International PhD Program central recruitment.

Recruitment process

We have set up a multi-step recruitment process with the goal of identifying people with right mix of skills, experience, and future potential. This involves an assessment of the application based on academic accomplishments and professional experience, presentations, structured interviews, and on-site visits .

Our selection process reflects the following structure:

The application requires details about your education (university, grades), research experience, interest and motivation,  an optional short video, degree transcripts and CV, as well as two reference letters, submitted independently by the referees. Please see detailed application instructions in the  application manual here .

 Applications via email can unfortunately not be accepted.

After submission of your application, you will receive an automatic email confirming that your application has been received. To be eligible for review, your application must be submitted before the application deadline.

Applicants will be chosen based on their academic qualification, aptitude, motivation, scientific interest and letters of recommendation.

Complete submissions will be forwarded to participating group leaders at the MDC and partner institutions.   Short-listed applicants will be invited for the 1st step of the recruitment process, online interviews. You can expect invitation letters 5-6 weeks after the application deadline.

Online interviews, held via videoconference platform, consist of of a welcome session, group leader presentation session, panel interviews and Q&A sessions with Grad Office, PhD Reps, and other interested PhD students and faculty members. 

Applicants participate in a 30-mins panel interview, including discussion on pre-recorded applicant's presentation, discussion on pre-selected scientific publication and general discussion. Candidates receive scores based on their  maturity and knowledge of the field, general knowledge, ability to express motivation for the PhD program, and clear interest in a specific research direction.

Candidates are ranked according to the panel scores, and the most successful candidates are invited to onsite interviews in Berlin. 

The days of onsite interviews in Berlin are dedicated to one-on-one interviews with group leaders, lab visits, and social events. 

The one-on-one interviews and lab visits provide sufficient amount of time for interviews, discussions, introduction to the lab and facilities, demonstration of working environment and dynamics, and interaction of group leader, candidate and lab members.

Evening social events are providing additional networking opportunities of candidates, PhD Representatives, lab members and faculty. Campus and city tours give candidates an opportunity for getting a better impression of their potential living and working environment.

Travel, visa expenses, accommodation and meals during interview days are fully covered by the MDC.  The   MDC cannot buy your transportation tickets in advance or reimburse the travel expenses before the Interview week.  Initially, you have to cover all of the travel expenses by yourself and the money will be reimbursed to you after the Interview week. 

On the final day of the interview week, candidates provide names of the research group(s) they would like to join for a PhD project. In a final closed plenary session for faculty, the candidates who will be accepted to the Int. PhD Program are determined. Results of the plenary session are announced within 1 week. 

Successful candidates start their research project within 4 months after the onsite interviews.

All candidates will receive feedback within a few days from the onsite interview week. Successful candidates will receive a position offer at this stage.

Transparency of the recruitment process

We value diversity and are committed to carrying out a transparent recruitment process for every position.

Our communication policy aims to maintain an objective and fair hiring process for all applicants. Communication between candidates and institute will be primarily conducted by e-mail. We acknowledge every submitted application immediately with an automatic e-mail response. During the hiring process, all successful applicants will receive notifications on the next steps via e-mail. In due course, candidates who have not been short-listed will be informed by e-mail. 

We adhere to principles of responsible research, good scientific practice, openness and transparency, and are committed to setting a good example as a scientific environment where scientists can flourish and grow, independently of their national or ethnic background, functional variation, sex/ gender identity/ alignment/ orientation, family configuration, or other such feature of their persons or contexts. 

Application manual

Once you have  registered on the Application portal , you will receive an e-mail with your personal login information. If this e-mail has not arrived, please check your spam folder. If the problem persists, contact us at  [email protected] . Please note that for the central recruitment we do not accept applications other than those submitted through our Application portal (e.g. CVs sent directly via e-mail or through other job application portals).

After registration, you may start working on your application, editing and saving the information in each section until you submit. The application should be completed in English. All mandatory fields are marked with *. Once you have submitted the form, you will no longer be able to edit the information, however, you can check the status of your recommendation letters. Please note that you will receive a confirmation email upon submission.

Recruitment Live Info Session

https://mdc-berlin.zoom.us/meeting/register/tJcvf-qorz0pE9RfFKS0NPmj-2nv5iB7uByF

Available projects.  After registration, the list of group leaders participating in this call and detailed description of their respective research projects can be found under the "Participating Group Leaders" tab. For more information on the participating project leaders please visit the  profile pages of our research labs . Carefully look into the research information and find projects that best fit your interests. Consider to check this list regularly during the call as more projects may be added.  Make sure to include all the labs and projects of your interest in the text of your application (see below, “Research & Motivation” section), with detailed statement of interest for each research lab.

Before starting the application process, please make sure that you are  eligible  to apply for our program. 

Below, you will find a detailed manual to help you fill in your application.  Please read it carefully, since incomplete and/or incorrectly filled applications will not be taken into further consideration.  Please make sure to include all relevant information in the application form. Information only included in additional documents (e.g. CV or certificates) might not be considered.

More information about eligibility, application process, language certificates, selection process, salaries, etc. can be found in our  FAQ  section.

Please note: it's applicant's responsibility to make sure the application, as well as the reference letters, are submitted before the deadline. Late submissions, incorrectly filled applications, and applications without references will not be taken into consideration. 

Personal Data: If you have multiple names, please enter the name from your passport. This is especially important if you need a visa to attend the onsite interview week in Berlin, since we will use it later for your visa application. If applicable, please use hyphens. Please enter the address that is available to and used by courier delivery services.  Make sure to use an address where you will be reachable in the period until the onsite interview week in Berlin  (especially at the time of visa applications, ca. 4-5 weeks before the onsite interview week). Furthermore, make sure to provide at least one telephone number. Additional Information:  Please tell us where you have first heard about the International PhD program at the MDC.

For detailed information on degree requirements, please check the “ Eligibility ” section. 

To be eligible, you need to have some research experience (bachelor and/or master research projects). Also, a research master thesis should have been required for obtaining your degree. If your study programs were structured in this way, you are eligible to apply.

Enter information about your university (both BSc and MSc). Please note that information on  both BSc and MSc  (or integrated program) need to be given, or you won’t be able to submit the application form.

Date of graduation:  If you have not graduated yet, put an approximate date of graduation. 

Final score:  To transform your grade into the German grading system, a detailed description of your university’s grading system is needed. Therefore, please provide information on i) highest possible grade in your grading system; ii) lowest passing grade in your system (please note: not possible, but passing) and iii) your achieved final grade. These will be used to translate your final grade into the German grading system, using the  Modified Bavarian Formula Calculator .

Scholarships and/or Awards:  Please clearly state the type of scholarships/awards you have received, the funding body, and the duration of the support.

Study abroad : Please state all long term (e.g. master program or a semester abroad), as well as short term (e.g. summer school, workshop or a course) visits. Please clearly state type of the program / event, host institution and duration of the study stay.

Research Interest: Please select the  research area  you are most interested in from the drop-down menu. In addition, you may also select drop-down research  key words  that particularly match your interests. If you would like to be considered for one of  the international exchange programs   at the MDC, select the appropriate program field from the drop-down menu. At this stage, this only indicates your interest, and it is not considered an application for the specific exchange program.  Research Experience : Please provide information on your relevant theoretical or experimental research experience, and list any supervised research projects with a duration of at least several weeks (BSc, MSc, internships, study visits or other). Please indicate whether your contribution to the project work was of a theoretical or experimental nature. Describe your role in the project, as well as contributions and results obtained from your work. Describe your accomplishments, with emphasis on the techniques you used and skills you obtained.  Publications & Conference  contributions : Please give a list of your publications ( only those published in international journals ) and conference contributions (poster presentation and/or talk). Please give full citations of all publications / conference contributions. Publications in preparation or under review should not be given. Research Statement - Motivation letter:  Use the research interest statement (Motivation letter) to express which  research field  you want to pursue during your PhD and why, as well as specify your  preferences for particular MDC research labs and projects .  Make sure to include all labs and projects of your interest, with detailed statement of interest for each project.  This might include, for example, the background of your interest (such as previous experience or personal motivation), details on scientific questions, concepts and/or techniques of interest, as well as your goals and vision of your scientific career development, etc. Here, you should also express and elaborate on any interest in an exchange program. Research Statement - Why did you select MDC for your application?  max. 2000 characters including spaces. Research Statement - Scientific discovery that has impressed you the most  (max. 2000 characters including spaces): Here, you should describe a scientific discovery that fascinates you, avoiding historical and technical overviews, but elaborating why you think this research matters. Did it disprove an existing paradigm, how did it affect everyday lives, or benefit the progress of society and science in general? Research Statement - Undergraduate courses you found the most inspiring:  max. 2500 characters including spaces.

Please enter complete contacts for  two referees . We recommend choosing referees that know you personally and are willing and able to write a description of your academic performance. After you fill in the contact details of your referees, an automated e-mail with a request for writing the recommendation letter will be sent to them. Please fill in  all required fields  - only after all fields are filled, the automated request will be sent. Please let  your referees know in advance that they will receive an invitation e-mail from MDC ! You cannot change your referee yourself once you have saved the contact. If you wish to change a referee, contact us in time to reset the information. Following the invitation, referees should submit their letters directly to the MDC PhD application system, using a link they received in the e-mail . Please make sure your referees have received an invitation email.  If they do not receive an e-mail within 24 hours after the automated request is sent, notify us at  [email protected]  immediately. 

Also, make sure that your referees have submitted their reference letters in time.  After submission, the status of your references will be indicated by the symbols on the left. The symbol will change from orange envelope to green "check mark" after your referees have submitted their letters.  Please note: References arriving after the reference deadline will not be processed and  applications will be considered incomplete . 

In this section you may upload a video where you address the following question in ca 2 minutes (min 1 minute):  "Which of the Nobel prize awards in the field of Physiology or Medicine, Chemistry or Physics did you find the most inspiring and why?" . You may elaborate on any award of the above categories, either recent or older.

All documents must be uploaded as pdf files (max of 3MB). 

A scanned version of the following documents is mandatory:

• Short CV  of max. 2 pages (In the CV, please provide only biographical data and professional development timeline. All relevant education, research experience, motivation, workshops, conferences and publications information should be given in the application form.)
• Transcript  (record of study, including courses list and scores)  of your highest degree awarded
• Certificate of your highest degree awarded

If you have not completed your Master's program yet, provide temporary transcripts, including scores. All documents  should be available in German or English . If you do not have an English version issued by your university, a translation certified by a public notary or other official will be sufficient. If your university does not provide transcripts, a marksheet should be provided instead.  Applications without scored transcripts cannot be further processed. 

A scanned version of the following documents is recommended ( but not obligatory! ):

• English proficiency test ​​​​​

Please note: Applicants  need to have a good command of written and spoken English , since the program language is English. Language proficiency tests for non-native speakers are recommended, however,  the proficiency certificates are not a requirement for the application !

Don’t forget to submit your application once you have completed all sections. After submission you cannot change or add information, but you can check the status of your references. The completed application must be submitted by the application deadline. 

Please note that upon submission you will receive a confirmation e-mail.

Register at our application portal

https://mdcphd.mdc-berlin.de/

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All PhD News

Humboldt-Universität zu Berlin

Over 200 years old,  Humboldt-Universität zu Berlin is a home to more than 35 000 students, with ca 16% of students coming from abroad.  Faculty of Life Sciences – Department of Biology of Humboldt-Universität is a long-standing partner of the MDC. In collaboration with MDC and Charité, Humboldt-Universität founded the Integrative Research Institute (IRI) for the Life Sciences in 2013. It pursues cutting-edge biomedical research with an interdisciplinary and inter-institutional approach.  

Charité Universitätsmedizin, Berlin

Charité Universitätsmedizin  is a joint medical faculty of the Humboldt-Universität zu Berlin and the Freie Universität, combining basic with clinical research and patient care. It is the largest university hospital in Europe, with more than 100 clinics and institutes. The MDC has a close partnership with the Charité: there are joint clinical research groups and major institutional collaborations.  

ECRC - Experimental and Clinical Research Center, Berlin

Established by the MDC and the Charité, within its 19 research labs and 2 facilities, the Experimental and Clinical Research Center (ECRC) integrates committed clinician/scientists and translational researchers in the research-driven environment on the MDC Campus in Berlin-Buch, offering excellent opportunities for education in translational research and training of the clinician scientist.  

Weizmann Institute of Science; Rehovot, Israel

The Weizmann Institute of Science is one of the leading multidisciplinary basic research institutions in the world, covering a broad range of studies in mathematics and computer science, physics, chemistry, biochemistry and biology. The Institute’s unique structure encourages cooperation among scientists from different disciplines, aimed at achieving the multi-disciplinarity necessary to provide innovative approaches.

The campus community currently numbers approximately 3.500, including 250 full-professors, associate professors and senior scientists who serve as heads of research groups, as well as 850 staff scientists, engineers and technicians, approximately 1.100 graduate students working toward MSc or PhD degrees and about 400 postdoctoral fellows.

In 2020, the MDC joined forces with the Weizmann Institute of Science, Humboldt-Universität zu Berlin and Charité Universitätsmedizin, starting  iNAMES - MDC-Weizmann Helmholtz International Research School (HIRS) for Imaging and Data Science from the NAno to the MESo.

Frequently asked questions:

Which degrees are recognised?

Applicants are required to hold (or obtain within 4 months of the onsite interview week) a university degree  equivalent to a Master Degree (MSc) , including  a research thesis , in  biology, biotechnology, medicine, molecular medicine, chemistry, computer science, data science, engineering, mathematics, physics, or some related field. 

Can I apply before finishing my master studies?

At the time of application, you do not have to have finished your degree. However, students are asked to  start their research project within 4 months of the onsite interviews , by which time you should anticipate to hold your degree.

I have a medical degree (MD). Am I eligible?

Yes, you are welcome to apply, as long as you have some relevant r esearch experience.

I have a degree in Public Health? Am I eligible?

If your master (or equivalent) program required a research project thesis to complete the program, AND you do have some relevant research experience in biology, biotechnology, medicine, molecular medicine, chemistry, computer science, data science, engineering, mathematics, physics, or some related field, you are eligible to apply. However, please note that the MDC is a research center, with a focus on basic biomedical research, and available projects are mostly not compatible with a Public Health background.  

My degree is from a university of applied sciences (Fachhochschule). Can I apply?

Yes, the graduates from the universities of applied sciences are welcome to apply if their university program required a research project thesis to complete the program, and they have relevant research experience.  

I only have a BSc, but a lot of research experience. Can I apply?

Unfortunately, neither a BSc degree alone, even of a 4-year-long program, nor additional experience represent an equivalent to the Master degree, thus you are not eligible. 

I already have a PhD. Can I apply for another?

If you  already hold a PhD degree, you are not eligible to apply .

Am I allowed to apply twice / again?

If you were not shortlisted for the online interviews, you are allowed to apply again. However, if you have already participated in the online interviews, you are not eligible to apply again. 

Does the program have language requirements and what is the program language?

The program language is English, therefore proficiency in English is a must. For your application, proof of English knowledge is recommended, but not mandatory!

Is there an age limit?

No, there is no age limit. However, please note that you should have obtained your MSc degree  (or equivalent) degree not more than four years prior to the interviews.

Where and when can I apply?

Applications for the Int. PhD Program are accepted  through the online Application portal only .  The portal opens twice a year, usually around May for Fall recruitment and in November for Spring recruitment. Exact dates and application details are indicated  here . 

Where can I find the list of available projects?

After registration, the list of group leaders participating in a call and detailed description of their respective research projects can be found under the "Participating Group Leaders" tab in the Application portal. For more information on the participating project leaders, please visit the profile pages of our research labs .

Do I need to select a specific project immediately?

You are not applying for a specific project at this stage, however, make sure to include all the labs and projects of your interest in the text of your application (“Research and Motivation” section), with detailed statement of interest for each research lab.

Can I save the data of my application and resume, completing it later?

Yes, you can save your application and take up editing again later. Once you have submitted your application, no more changes can be made. 

What information has to be submitted with the application?

You will submit information on education, research skills and interests, a letter of motivation and contacts of 2 referees. In the document section, you are asked to upload a CV, university transcripts and certificates, and if available, the certificate of an English language test. English language tests, GRE, etc are not mandatory at this stage. For details, visit the application manual.

What happens if I cannot provide any English language certificates?

Proof of English language knowledge is not obligatory but rather  optional . If you do not have such documents, your application will still be valid.

Do I need to have my certificates available in English?

All documents should be available in German or English. If you do not have an English version issued by your university, a translation certified by a public notary or other official will be sufficient.

My PDF files are too big. What can I do?

You will have to reduce their sizes. There are several online free tools that can easily reduce the size of your PDF files, for example:

• http://smallpdf.com/compress-pdf
• https://www.pdfcompress.com/

Please make sure you understand their privacy policy before using any of these third-party services.

How many recommendation letters do I need to have and can I send the PDFs (scans) directly?

You will need recommendation from  two referees . We recommend choosing referees that know you personally and are willing and able to write a description of your academic performance. As soon as you have completed the contact details, your referees will automatically receive a request for a letter of recommendation and they should submit their letters directly using a link they receive in the e-mail. Thus, you should not send the PDFs of the recommendation letters directly to us.

My referee did not receive an invitation from the MDC to send a recommendation letter, what should I do?

Firstly, please make sure you entered the correct e-mail address of your referee. Secondly, your referee should check their spam mailbox to make sure the invitation letter did not end up there. If you entered the wrong address or the letter simply did not arrive, contact us immediately at  [email protected]

Will I receive a confirmation after having registered and submitted my application?

Yes, after having registered you will receive a confirmation email to the mail address stated. After you have submitted your application you will again receive a confirmation. If you did not receive any confirmation email, most likely our email ended up in the spam or junk folder. If you cannot find your confirmation email there, you might have misspelled your email address during registration. In that case, simply fill in the registration form again. If you still do not receive your registration email, please contact the graduate school office [email protected] .

What happens after submission?

Applications are prescreened and the MDC faculty selects candidates for the first step of the recruitment process, online interviews. Invitations for online interviews will be sent about 5-6 weeks after the application deadline.

How can I check the status of my application?

Once you have submitted the form, you will no longer be able to edit the information, however, you can check the status of your recommendation letters, using the colour code of the box. The symbol on the left will change from an orange envelope to a green "check mark" after your referees have submitted their letters. Other information about changes in the status of your application (submission, invitation / rejection decision) will be communicated via e-mail.

How and when will I hear about the outcome of my application?

Students who submitted a complete application will be informed by email. Students shortlisted for online interviews will be informed about the result 5-6 weeks after the application deadline. All other candidates will be informed shortly after.

What happens during the online interviews?

Applicants participate in a 30-mins panel interview, including discussion on pre-recorded applicant's presentation, discussion on pre-selected scientific publication and general discussion. Candidates receive scores based on their maturity and knowledge of the field, general knowledge, ability to express motivation for the PhD program, and clear interest in a specific research direction.

How should I present my research project?

Candidates present their research project (usually master research project or other longer project they participated in during their studies) in form of a pre-recorded presentation. Presentations will be stored in the application portal and available to all group leaders for viewing before the start of online interviews.

Panel members will watch the pre-recorded candidate presentations before the start of online interviews and prepare questions for the short presentation discussion.

How should I prepare for the online interviews?

You will receive all the instructions and technical details about the videoconference platform used for online interviews well in advance. The most important thing is to get familiar with the videoconference system, and get comfortable with its functions and features, such as navigation between rooms or screen sharing. Second important point is to make sure your internet connection is fast enough to support this kind of activity. Here, it is better to use the ethernet cable connection instead of WiFi. Please check the settings and quality of both audio and video features. If your home internet connection is not fast enough, you should find a suitable alternative, such as using resources at your university or public library. If possible, you should also prepare a second screen, since this eases videoconference participation.

What if problems with the videoconference system arise during my online interviews?

We will prepare more than one videoconference platform for you to use, and backup communication channels will be available. 

When will I know if I got selected for the onsite interviews in Berlin?

You will receive invitations for the onsite interview week in Berlin within 1 week of the online interviews.

What happens during the onsite interview week?

Onsite interview week consist of introduction to the Grad School program, one-on-one interviews, lab visits, campus and city tours and social events.

The one-on-one interviews and lab visits provide sufficient amount of time for interviews, discussions, introduction to the lab and facilities, demonstration of working environment and dynamics, and interaction of group leader, candidate and lab members.

Evening social events are providing additional networking opportunities of candidates, PhD Reps, lab members and faculty. Campus and city tours give candidates an opportunity for getting a better impression of their potential living and working environment.

Will interview costs be covered?

All costs during interview week including travel expenses, visa, accommodation and food will be covered by the MDC. Please note:  MDC as a public institution is not able to buy your transportation tickets in advance or reimburse the travel expenses before the Interview week.  Initially, you have to cover all of the travel expenses by yourself and the money will be reimbursed to you after the Interview week. 

What happens if I cannot participate in the Interview week in Berlin due to time constraints?

Your participation is absolutely obligatory. If you are not available on that week, you would unfortunately not be considered for the selection process. Videoconference options are not possible at this stage.

When do I know whether I have been accepted to the program?

Acceptance letters are sent within 7-10 days after interviews.

When can I start my PhD?

Students have to start their research project  within 4 months of the onsite interviews  at the MDC.   In general, PhD students will agree with his or her prospective supervisor on a specific starting date.

How do I enroll at the university?

Upon your start at the MDC, the MDC Grad School Office will lead you through the process of the enrolment. Most students at MDC are enrolled at the local partner universities Charité, Humboldt Universität, or Freie Universität. Usually, y ou will enroll at the institution your prospective group leader is affiliated with. 

Are there any tuition fees?

There are no MDC Grad School tuition fees, however, after enrolling at the university you shall pay the semester fees of ca. 350 Euro, including public transportation ticket.

How  will I finance my PhD studies ?

PhD students receive a work contract according to the German public system, including health and social insurance, as well as pension scheme. The initial contract is for 3 years and the net salary amounts to about 1600 Euro per month.

Is there support upon arriving in Berlin?

The Welcome and family office at the MDC supports all students upon arriving in their first steps settling in Berlin,  including all issues related to VISA and residence.

How long does it take PhD students to complete their PhD?

Students are expected to complete their degree in four years.

Contact and information

[email protected]

Graduate School

Application portal

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MD/PhD program Molecular Medicine

Fully accreditated in 2009 as the first PhD program in Germany according to European standards (ECTS), the MD/PhD program features an interdisciplinary curriculum leading to a PhD thesis after 3-4 years.

The program is a joint venture between the following institutions :

• Hannover Medical School
• Leibniz University Hannover
• Fraunhofer Institute ITEM Hannover
• Helmholtz Centre for Infection Research (HZI) Braunschweig
• TWINCORE (Hannover)
• NIFE (Hannover)

PhD topics cover a broad spectrum, including the following areas :

• Immunology & Infection
• Molecular & Cell Biology
• Hematology & Oncology
• Biochemistry & Structural Biology
• Stem Cell Research

and many more....(nearly all clinics / institutes doing molecular experimental research are involved ☺)

Faculty : more than 150 scientists ( details )

Potential PhD students : Holders of an M.Sc. or equivalent degree in Medicine, Biology, Biochemistry, Biotechnology, Biomedicine etc.

Degree:   you will either receive a PhD or Dr.rer.nat. degree (later only for life scientists)

Medical students/graduates are invited to check MD/PhD education for more clinical training options. Please note that we only award a PhD title (not an "MD/PhD")

What we offer : Interesting projects, in-depth basic science seminars for the first two years, excellent structure and supervision. 

Next application deadline : March 1st, 2024. Online application from December 1st 2023 at

https://hbrs.cloud.opencampus.net/

More information

Structured Doctoral Programs

Each of our structured doctoral programs offers a comprehensive and cross-disciplinary curriculum designed to help you realize your full potential and prepare for a successful career. The programs include innovative, personalized advising with regular progress checks, as well as extensive opportunities to broaden your research network and connect with peers in your field.

The University of Bonn offers a wide range of funding opportunities, which have been summarized for you on this page, divided into the following categories:

Bonn International Graduate Schools (BIGS)

Phd programs within our cluster of excellence.

• Structured Doctoral Programs by Discipline 

Third-Party Funded Programs

Doctoral education at the highest level: BIGS enable doctoral studies in outstanding research contexts with attractive international collaborations and a qualification program tailored to the needs of graduate students.

Located at the Hausdorff Center for Mathematics and supported by Germany’s Excellence Initiative, BIGS-M  is home to all of the University’s doctoral candidates in mathematics and contributes to Bonn’s excellent international reputation in the field.

BGSE offers a structured program that is tailored to the needs of doctoral candidates, including an internationally recognized research network.

Supported by Germany’s Excellence Initiative and jointly administered by the renowned Physics Institutes at the Universities of Bonn and Cologne, BCGS  offers doctoral studies through an integrated honors program.

Home to an international community of talented biomedical scientists, BIGS DrugS 6 6 is the hub for doctoral candidates from pharma research institutes within the University’s Faculty of Mathematics and Natural Sciences and Faculty of Medicine.

BIGS-OAS offers a wide range of courses within a research context, focused on the cultures and societies of Asia and Asia Minor.

BIGS Neuroscience provides a top-level, internationally competitive program in this rapidly growing field.

BIGS CPS's interdisciplinary approach combines medical, agricultural and pharmaceutical research.

BIGS Chemistry 10 doctoral candidates enjoy an exceptional and ambitious program covering all fields of chemistry.

This three-year doctoral program  is offered in conjunction with the University’s ImmunoSensation Cluster, which is funded by Germany’s Excellence Initiative.

Part of the University of Bonn’s Center for Development Research, BIGS-DR   trains students for an international career in development cooperation, policy or research through a combination of academic study and intensive tutorship.

The BIGS Land and Food combines the research at the agricultural Faculty with an interdisciplinary study program.

Clusters of Excellence stand for international and interdisciplinary elite research and offer young scientists excellent funding and career conditions. The University of Bonn currently has six clusters of excellence, more than any other university in Germany, and thus opens up a broad spectrum of possible research topics to doctoral candidates. Here you will find an overview of the university's clusters of excellence.

PhD Programs within our Excellence Cluster

The goal of the Hausdorff Center of Mathmatics is to identify and address mathematical challenges of the 21st century, to advance groundbreaking fundamental mathematical research worldwide, and to develop the mathematical methods and tools required by science and society.

Part of the Hausdorff Center is also a graduate school: The Bonn International Graduate School of Mathematics (BIGS-M) hosts all doctoral students of mathematics and contributes to the outstanding international reputation of the university in this field. The duration of the program is usually 3 years, and the doctorate (Dr. rer nat.) can be earned as a degree.

More information: https://www.bigs-math.uni-bonn.de/de/studies/ 14 15 15

ImmunoSensation2 aims to continue the success story of the existing ImmunoSensation cluster. While the emphasis so far has been on fundamental research in particular of the innate immune system, now the mechanisms of immune intelligence are to be uncovered, i.e. the question of how the body succeeds in adapting immune responses to specific situations and then remembers this in order to be optimally prepared for similar challenges in the future. The cluster's graduate school, the Bonn International Graduate School Immunosciences and Infection offers a structured, three-year doctoral program.

You can find further information about this program here: 

https://www.immunosensation.de/opportunities/young-scientists

Until today, dependency studies has almost exclusively dealt with slavery on the American continent or in antiquity. The Cluster of Excellence "Bonn Center for Dependency and Slavery Studies (BCDSS)" aims to broaden this perspective in terms of content, space and time. Within the framework of the cluster, a structured doctoral program with a duration of 4 years is offered.

Further information can be found at:  https://www.dependency.uni-bonn.de/en 15 16

Over the last few decades, computer hardware has become smaller and smaller, but their technology remains more or less the same. Slowly, this development is reaching its limits.Thus, we need new technologies that satisfy our growing hunger for even more powerful hardware.

Quantum physics could be a solution.

Together with the University of Cologne and the RWTH Aachen, Bonn researchers want to work on making this new technology usable. To achieve this, quantum bits or even qubits - the quantum counterpart to our previous bits - quantum communication channels that build networks and error correction methods have to be explored from the ground up. As part of the Excellence Initiative, the Bonn-Cologne Graduate School of Physics and Astronomy (BCGS) offers a doctoral program with an integrated honors program.

Further information can be found at:  http://www.gradschool.physics.uni-bonn.de/. 4 4

The ECONtribute researches the functioning of markets as well as reasons for their failure. In doing so, the cluster goes beyond traditional analyses by systematically combining model-based theoretical approaches and behavioral explanatory models while incorporating legal and political frameworks. Within the cluster, the Bonn Graduate School of Economics (BGSE) offers doctoral students a tailored structured doctoral program that includes an internationally recognized research network.

Further information can be found at:  https://www.bgse.uni-bonn.de. 3 3

Increasing agricultural production despite limited land while reducing the ecological footprint of agriculture - this is one of the challenges of our time. For this reason, the University of Bonn and Forschungszentrum Jülich are jointly developing methods and new technologies to observe, analyze, better understand and more specifically treat plants. The cluster's graduate school, the Theodor Brinkmann Graduate School, offers an interdisciplinary study program to master's students and doctoral candidates at the Faculty of Agriculture.

More information: https://www.phenorob.de/ .

The Third-Party Funded Programs at the University of Bonn offer structured doctoral studies on selected research topics. They enable close networking among doctoral students conducting research on related topics.

Bonn International Graduate School of Mathematics (BIGS-M) 2 17 18 18 Located at the Hausdorff Center for Mathematics, BIGS-M provides an umbrella for all Bonn PhD students in mathematics. Thus, the BIGS-M contributes to the excellent national and international reputation of mathematics at Bonn.

Bonn International Graduate School Immunosciences and Infection The BIGS Immunosciences and Infection is a structured 3-year PhD program in conjunction with the ImmunoSensation Cluster/Bonn. The ImmunoSensation Cluster is part of the Excellence Strategy.

DFG Research Training Group "Gegenwart/Literatur. Geschichte, Theorie und Praxeologie eines Verhältnisses" (GRK 2291) [only in German]

The Research Training group supported by the DFG  aims at the exploration and analysis of the constitutive dimensions of the concept of contemporary literature.  

DFG international Research Training Group "Myeloid antigen presenting cells and the induction of adaptive immunity" GRK (2168) 19 19 19 19 The DFG-funded project is a cooperation of the University of Bonn and the University of Melbourne. The principal research focus is the intersection between innate and adaptive immunity in the context of infection.

DFG Research Training Group  "The Macroeconomics of Inequality"  ( GRK 2281) 20 20 20 20 The research program focuses on the macroeconomic aspects of inequality, an aspect of first-order importance for society. 

DFG Research Training Group "Template-designed Organic Electronics (TIDE)" (GRK 2591) 21 21 21 The Graduate Program 'Template-Designed Optoelectronic Devices' (TIDE) aims to provide comprehensive doctoral education in the field of Organic Electronics (OE) to meet the requirements of highly qualified and multidisciplinary professionals. 

DFG Research Training Group "Tools and Drugs of the Future - Innovative Methods and New Modalities in Medicinal Chemistry" (GRK 2873)

The goal of the RTG " Tools and Drugs of the Future" is to modernize medicinal chemistry and train a new generation of medicinal chemists and researchers at the interface with interconnected disciplines. In addition, the projects are intended to contribute to the development of new drug substances.

Integrated Research Training Group at the DFG Collaborative Research Centre "Synaptic Micronetworks in Health and Disease" (SFB 1089) 22 22 27 27 Located at the newly inaugurated SFB 1089 on neuronal networks, the Integrated Research Training Group offers a structured graduate program for all doctoral researchers at the Centre.

Integrated Research Training Group at theDFG Collaborative Research Centre "Future Rural Africa" (SFB/TR 228) The integrated research group is investigating the relationship between land use change and shaping the future in rural africa in a total of 14 subprojects.

Integrated Research Training Group at the DFG Collaborative Research Centre "Open System Control of Atomic and Photonic Matter" (SFB/TR 185) 24 The collaborative research centre Oscar will explore the physics of open systems.

Integrated Research Training Group at the DFG Collaborative Research Centre "Aortic Diseases" (SFB/TR 259) 25 The aim of this research initiative is to better understand the molecular and cellular mechanisms of resident and non-resident cells in aortic diseases.

Integrated Research Training Group at the DFG Collaborative Research Centre "Regional Climate Change: Disentangling the Role of Land Use and Water Management" (SFB 1502) The SFB combines the strengths of the University of Bonn and its project partners to answer one of the most difficult questions in understanding climate change.

Integrated Research Training Group at the DFG Collaborative Research Centre "Brown and Beige Fat - Organ Crosstalk, Signaling and Energetics (BATenergy)" (SFB/TRR 333) The CRC investigates metabolism/diabetes and focusses on brown adipose tissue. 

One Health and Urban Transformation

The NRW Forschungskolleg One Health and Urban Transformation is a transdisciplinary graduate school that aims to find interventions to achieve optimal health for humans, animals, plants and the environment with a special focus on developments in NRW, Saõ Paulo, Accra and Ahmedabad.

International Max Planck Research School Moduli Spaces 27 27 In cooperation with the University of Bonn, the renowned Bonn Max-Planck-Institute for Mathematics offers a PhD program with a special focus on the study of moduli.

International Max Planck Research School for Astronomy and Astrophysics 28 28 In cooperation between the Max-Planck Institute for Radio Astronomy and the Universities of Bonn and of Cologne, the Research School facilitates 3 years of PhD studies with a curriculum tailored to the individual student.

International Max Planck Research School for Brain and Behavior 29 The IMPRS for Brain & Behavior is a cooperation between the Max Planck Institute for Neurobiology of Behavior - caesar, the University of Bonn and the German Center for Neurodegenerative Diseases (DZNE) in Bonn

International Max Planck Research School - Recharge IMPRS-RECHARGE focuses on interdisciplinary research between chemistry and physics with an emphasis on catalytic mechanisms, physical-chemical analysis and energy topics. Scientific challenges shall be looked at from different angles. Furthermore the combination of theory and practice is a vital aim of the IMPRS-RECHARGE.

Marie Curie Innovative Training Network "Macro and Microplastic in Agricultural Soil Systems“ (SOPLAS) The SOPLAS project will assemble a multidisciplinary team to study the nexus of plastic–agriculture–soil. It will also train a new generation of leading experts. The project aims to identify the plastic cycle within agricultural soil systems and support the development of environmental policies related to mitigating the impact of plastics. The findings will advance our knowledge about the sustainable use of plastics in European agriculture.

Marie Curie Innovative Training Network "Early Stage Researchers EDUCational Program on Factor VIII Immunogenicity“ (EDUC8 ) 32 37 The EDUC8 program is a multidisciplinary training program with exposure of the enrolled ESRs to a core common educational package and development of individual PhD researchprojects dedicated to decreasing the societal burden associated with the development of anti-FVIII antibodies in Europe.

Marie Curie Innovative Training Network "Research and Training in Early Life Nutrition to Prevent Disease" (GROWTH)

GROWTH is an Innovative Training Network that aims to train young business-oriented researchers in developing pathological insights, biomarker diagnostics and personalized nutritional interventions for intestinal failure in neonates and preterm infants.

Tools4Teams - "Research Training to Design and Implement Tools Supporting Safe Teamwork in Healthcare"

The Tools4Teams research project will prepare the next generation of teamwork experts to contribute new insights and smart technologies for safe and effective care. Tools4Teams brings together expertise from social and technical sciences, human-centered design, education, and clinical specialties.

Trinational Graduate College "Mass and Integration in Antique Societies" [in German/French] Supported by the Deutsch-Französische Hochschule since 2011, the tri-national Graduate School in Ancient History offers curriculum events in Bonn, Berne, and Strasbourg.

Structured Doctoral Programs by Discipline

Find the right structured doctoral program at the University of Bonn in your discipline here:

• Cross-Disciplinary Options
• Medicine and Life Sciences
• Mathematics and Natural Sciences
• Agriculture

Faculties at the University of Bonn work together to design interdisciplinary programs that combine key perspectives and offer unique insights.

Cross-Disciplinary Programs

Bonn International Graduate School for Development Research (BIGS-DR) 42 Unique in Europe, BIGS-DR links perspectives from the Faculties of Philosophy, Agriculture, and Law and Economics – with an international focus.

Bonn International Graduate School of Neuroscience (BIGS Neuroscience) 8 8 A collaboration between the University’s Faculty of Medicine and Faculty of Mathematics and Natural Sciences, as well as external partners, BIGS Neuroscience offers a medical program alongside five research areas in medicine.

SciMed Doctoral College 43 42 The Doctoral College offers scientific training for students in medicine and dental medicine, leading to a dual Dr. med. and Dr. med. dent. degree.

Researchers at the University of Bonn explore a wide variety of issues in economics, including game theory, applied microeconomics, monetary and international macroeconomics, contract theory, labor economics and finance.

Economics Programs

Bonn Graduate School of Economics (BGSE) BGSE offers a structured program that is tailored to the needs of doctoral candidates, including an internationally recognized research network.

DFG Research Training Group "Die Macroeconomics of Inequality" (GRK 2281) The research program focuses on the macroeconomic aspects of inequality, an aspect of first-order importance for society. 

Law Programs

Graduate School of Law and Political Science Department of Law The Graduate School of the Faculty of Law and Political Science was founded in the summer semester of 2018 and supports the doctoral students in preparing their doctoral studies.

The University of Bonn’s Faculty of Medicine offers doctoral programs in medical biochemistry, neurosciences and pharmacology. With the exception of the SciMed Doctoral College, all programs are administered in cooperation with the University’s Faculty of Mathematics and Natural Sciences.

Cross-Disciplinary Program

SciMed Doctoral College The Doctoral College offers scientific training for students in medicine and dental medicine, leading to a dual Dr. med. and Dr. med. dent. degree.

Neuroscience

Bonn International Graduate School of Neuroscience (BIGS Neuroscience) BIGS Neuroscience provides a top-level, internationally competitive program in this rapidly growing field.

Synaptic Micronetworks in Health and Disease (SFB 1089) Supported by the German Research Foundation (Deutsche Forschungsgemeinschaft – DFG) collaborative research centers, this integrated research training group works to identify fundamental rules that govern neuronal behavior at the network level and translate network dynamics to mammalian and human behavior.

International Max Planck Research School for Brain and Behavior A joint venture of the University of Bonn, the Max-Planck-associated Center of Advanced European Studies and Research, the Max Planck Florida Institute for Neuroscience, and Florida Atlantic University, this graduate school offers a complete doctoral and research program in the neurosciences.

Marie Curie Initial Training Network "modelling and pRedicting Human decision-making Using Measures of subconscious Brain processes through mixed reality interfaces and biOmetric signals" (RHUMBO) RHUMBO proposes using measures of subconscious brain processes through the use of mixed reality technologies (MRT) and advanced biometric signals processing as a new paradigm to improve the knowledge that implicit brain processes have in human decision-making.

Pharma Research

Bonn International Graduate School of Drug Sciences (BIGS DrugS) Home to an international community of talented biomedical scientists, BIGS DrugS is the hub for doctoral candidates from pharma research institutes within the University’s Faculty of Mathematics and Natural Sciences and Faculty of Medicine.

Pharmacology of 7TM-Receptors and Downstream Signaling Pathways (GRK 1873) Supported by DFG, this group combines expertise in the areas of pharmacology and pharmacy.

Bonn International Graduate School of Immunosciences and Infection

This three-year doctoral program is offered in conjunction with the University's ImmunoSensation Cluster , which is funded by Germany’s Excellence Initiative.

DFG Research Training Group "Myeloid antigen presenting cells and the induction of adaptive immunity" GRK (2168) The DFG-funded project is a cooperation of the University of Bonn and the University of Melbourne.

At the University of Bonn’s Faculty of Arts, you’ll find a highly international environment with students and researchers in a wide range of fields.

German Studies, Comparative Literature and Culture 

Structured Doctoral Program in German Studies (SPP) [website in German] Taught in German, the SPP supports doctoral candidates’ initiatives within the Institute for German, Comparative Literature and Cultural Studies.

German-Italian Doctoral College [website in German] Taught in German, this three-year grant program provides structured doctoral studies for researchers in German and Italian, with time in both Bonn and Florence.

History and Ancient History 

Mass and Integration in Antique Societies [website in German and French] Supported by Franco-German University and taught in German and French, this trinational doctoral program includes study in Bonn; Berne, Switzerland; and Strasbourg, France.

Oriental and Asian Studies Bonn

International Graduate School of Oriental and Asian Studies (BIGS-OAS) BIGS-OAS offers a wide range of courses within a research context, focused on the cultures and societies of Asia and Asia Minor.

Romance Studies

Italian Studies [website in German and Italian] Offered in cooperation with the Universities of Florence and Paris-Sorbonne IV, this trinational doctoral program is taught in German and Italian.

Structured DPhil program at the Faculty of Arts The program supports qualified doctoral candidates from all disciplines in their doctoral projects. It provides the opportunity for networking, interdisciplinary exchange in diverse social sciences and humanities subjects, progress monitoring and financial support for travel, workshops or research funding as part of the doctorate.

European Founding Myths in Literature, Arts and Music [website in German, French and Italian] This trinational program is jointly organized by the Universities of Bonn, Florence and Paris-Sorbonne IV and taught in German, French and Italian.

Bonn International Graduate School for Development Research (BIGS-DR) Part of the University of Bonn’s Center for Development Research, BIGS-DR trains students for an international career in development cooperation, policy or research through a combination of academic study and intensive tutorship.

The University’s Faculty of Mathematics and Natural Sciences offers numerous externally funded doctoral programs in areas including mathematics and informatics, physics, biology, pharmacology and molecular biomedicine.

Programs in neuroscience, pharma research, immunoscience, and infection and molecular biomedicine are offered in cooperation with the Faculty of Medicine.

Mathematics

Bonn International Graduate School of Mathematics (BIGS-M) 2 2 Located at the Hausdorff Center for Mathematics, BIGS-M is home to all of the University’s doctoral candidates in mathematics and contributes to Bonn’s excellent international reputation in the field.

International Max Planck Research School on Moduli Spaces 53 53 This program includes courses, seminars and activities focused on the geometric spaces whose points represent fixed algebro-geometric objects (or isomorphism classes of such objects).

Physics und Astronomy

Bonn-Cologne Graduate School of Physics and Astronomy (BCGS) 4 4 Supported by Germany’s Excellence Initiative and jointly administered by the renowned Physics Institutes at the Universities of Bonn and Cologne, BCGS offers doctoral studies through an integrated honors program.

International Max Planck Research School of Astronomy and Astrophysics 55 55 This program offers a broad spectrum of topics in observational and theoretical galactic and extragalactic astrophysics, observational and theoretical cosmology, and fundamental physics – using astronomical tools and instrumentation.

Leibniz Graduate School on Genomic Biodiversity Research Based at Bonn’s Alexander Koenig Research Museum, this school is focused primarily on insect genome evolution.

Bonn International Graduate School of Chemistry (BIGS Chemistry) 57 57 BIGS Chemistry offers an internationally competitive doctoral program and opportunities to perform cutting-edge research.

Neurosciences

Bonn International Graduate School of Neuroscience (BIGS Neuroscience) 8 8 BIGS Neuroscience provides a top-level, internationally competitive program in this rapidly growing field.

Synaptic Micronetworks in Health and Disease (SFB 1089) 22 22 Supported by DFG collaborative research centers, this integrated research training group works to identify fundamental rules that govern neuronal behavior at the network level and translate network dynamics to mammalian and human behavior.

International Max Planck Research School for Brain and Behavior 29 29 The IMPRS for Brain & Behavior is a cooperation between the Max Planck Institute for Neurobiology of Behavior - caesar, the University of Bonn and the German Center for Neurodegenerative Diseases (DZNE) in Bonn.

Bonn International Graduate School of Drug Sciences (BIGS DrugS) 6 6 Home to an international community of talented biomedical scientists, BIGS DrugS is the hub for doctoral candidates from pharma research institutes within the University’s Faculty of Mathematics and Natural Sciences and Faculty of Medicine.

Pharmacology of 7TM-Receptors and Downstream Signaling Pathways (GRK 1873) 18 18 Supported by DFG, this group combines expertise in the areas of pharmacology and pharmacy.

BIGS Immunoscience and Infection A structured, three-year doctoral program, IITB is offered in conjunction with the ImmunoSensation Cluster at the University of Bonn.

Doctoral candidates in the field of agriculture may choose to study through the Faculty of Agriculture’s Theodor Brinkmann Graduate School or earn their degree through the University of Bonn’s Center for Development Research.

Agriculture Programs

Bonn International Graduate School for Land and Food (BIGS Land and Food)  Founded in 2008, the Brinkmann School is home to master's and doctoral candidates in the Faculty of Agriculture, combining research with an interdisciplinary study program.

Bonn International Graduate School for Development Research (BIGS-DR) 12 Part of the University of Bonn’s Center for Development Research 59 , BIGS-DR trains researchers for an international career in development cooperation, policy or research through a combination of academic study and intensive tutorship.

Dr. Robert Radu

[email protected]

+49 228 73-60222

Poppelsdorfer Allee 47

Office Hours

Questions about the structured doctorate? Register for the (virtual) office hours and get advice:

• Tuesday 2.00 p.m. - 4.00 p.m.

Additional Qualification: Doctorate plus

Expand your skills with our training program Doctorate plus.

Learn about the numerous funding opportunities and grants for doctoral students.

Events and Opportunities

Find out what's new and see upcoming events.

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Jobs & Career

International phd program seeing beyond..

The DKFZ International PhD Program in Heidelberg (also known as the Helmholtz International Graduate School for Cancer Research) is the interdisciplinary structured graduate school for all PhD students working at the German Cancer Research Center (DKFZ) . Our PhD students conduct research at the forefront of basic, computational, epidemiological and translational cancer research.

The application deadline is 24 April 2025 .

Click here for more information on the selection process and how to apply.

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“When I found out about the international PhD Program at the DKFZ, I knew this was where I wanted to be: Cutting edge cancer research in one of the most renowned institutes in Germany in a beautiful and multicultural city is the perfect combination. My PhD studies at the DKFZ are a unique experience not only academically but they also enable my personal growth and cultural exchange.” (Antonino from Argentina, PhD student at the DKFZ)

You are thinking of doing your doctorate in Germany ? Then have a look at this Guide to doing a PhD in Germany which we wrote in cooperation with jobs.ac.uk and at this video by Research in Germany .

If you are a bachelor student or you've just started your master's studies , please take a look at our programs for undergraduate students !

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PhD study program

The PhD program in Molecular Medicine aims at an in-depth education in the translation of basic knowledge within molecular biosciences into applications regarding the prevention, diagnostics and therapy of human diseases. The Faculty of Medicine at the University of Göttingen has a long tradition and is further expanding its programs towards such research goals The establishment of a PhD program in Molecular Medicine represents only the most recent effort of this kind. The program is part of the Georg August University School of Sciences (GAUSS) and the Göttingen Graduate School for Neurosciences, Biophysics, and Molecular Biosciences (GGNB) .  

The program is aiming at the education of PhD students in these disciplines and technologies. The candidates are expected to conduct a high-profile research project in one of the afore-mentioned subjects. They will be supported by stringent supervision, thesis committees, methods courses and informal counseling. Overall, the program aims at the thorough education of young scientists who address medically relevant problems with a sound background in basic sciences.

PhD Program in Experimental Medicine

Since the winter semester 2012/2013, the Medical Faculty of the University of Tübingen has been offering a PhD program in Experimental Medicine. The target group are graduates of medical courses (human medicine , dentistry ) and related subjects such as molecular medicine medical technology , natural sciences , who are interested in an intensive scientific education and would like to prepare for work in the field of medical research and basic research After successful completion of the PhD program , the academic degree of Doctor of Philosophy" (PhD) in Experimental Medicine is awarded by the Medical Faculty of the University of Tübingen.

The PhD Program

The phd program in a nutshell.

To strengthen the link between science and applied medicine by preparing and equipping excellent students for a career in life science

To provide excellent young researchers in medicine and the life sciences with advanced professional qualifications for a career in research

REQUIRED DEGREE

Degree in medicine or a Master of Science in the life sciences or related disciplines

RESEARCH AREA

All areas of biomedical and clinical research , with a focus on infection medicine and microbiology immunology, oncology , neurosciences , pharmacology , cardiology , diabetology , imaging science , biomedical health services research , epidemiology , clinical research and public health

PhD in Experimental Medicine or MD/ PhD in Experimental Medicine double degree

TUITION FEES

Winter semester (October) or summer semester (April)

PROGRAM DURATION

Six semesters (three years), with a possibility to combine the program with medical studies and/or medical residency

Funding is mostly arranged by the student (applying for scholarships) or the research group (through project funding). Useful information on funding a PhD study in Germany is supplied by the German Academic Exchange Service (DAAD).

Components of the PhD Program

The main component is the individual research project to be submitted in form of a doctoral thesis.

SUPERVISION 

During your PhD study, you are supervised by your doctoral committee a group of three excellent scientists from the Faculty of Medicine, the Faculty of Science or external research institutes.

Additional to the PhD research, an individual academic course program (lectures, seminars, laboratory training) is defined to equip you with useful knowledge and skills: You must acquire 30 ECTS consisting of

• 8 ECTS from the compulsory programme (Lecture series, Journal club and or progress reports, Good scientific Practice course)
• 12 ECTS from Practical offers and courses of the different modules included in our curriculum
• 10 ECTS to be chosen individually

Courses and practical offers for the PhD Program can be found in the event database of the talent academy

PhD Candidate Resources

The phd board.

The board evaluates specific inquiries in compliance with the PhD regulations Head of the board : Prof. Dr. Olga Garaschuk Deputy head of the board : Prof. Dr. med. Christian la Fougère Faculty board members with core module

• Prof. Dr. Michael Schindler, Immunology and Infection
• Prof. Dr. med. Sabine Belard, Infection and Tropical Medicine
• Prof. Alexander N. R. Weber, PhD, Immunology
• Prof. Dr. Sandra BeerHammer, Immunology and Pharmacology
• Prof. Dr. med. Markus Hahn, Oncology and Surgery
• Prof. Dr. med. Ulrich Lauer, Oncology
• Prof. Dr. Marlies Knipper, Neurology
• Prof. Dr. Daniela Thorwarth, Data Science
• Prof. Dr. Tassula Proikas Cezanne ( Representative Sciences Faculty), Molecular Biology
• Prof. Dr. Robert Lukowski (Deputy representative Sciences Faculty),  Pharmacology

PhD representatives

The PhD Representatives are three to four candidates selected by the PhD candidates for a period of one years. They are in close contact with the program coordinator and faculty and report any concerns, feedback, or ideas from the PhD candidates regarding all aspects of the program.

Coordination Office

The coordination office assists PhD candidates with inquiries and concerns regarding academic matters, program activities, course registrations, transcripts, and personal issues. In addition, the coordination office is in constant contact with the PhD Board and communicates decisions to candidates regarding new policies, admissions, and special inquiries.

Selection and admission

Application

Application and admission procedure.

A prerequisite for the application is the commitment of a university professor of the Medical Faculty Tübingen to supervise the doctorate and funding.

Important dates 

Application deadline

• for the summer semester 15.01.
• for the winter semester 15.06.
• upon agreement

Only complete applications that are received by the deadline of the respective semester and meet the admission requirements can be considered in the selection process. The exact application requirements can be found in the "Application Guidelines" (download area). The competition is divided into two stages: The evaluation of the written application and the personal interview.

The written application

The application must include an exposé for the planned research project and the confirmation of care of a habilitated member of the Faculty of Medicine.

Information on possible research projects can be obtained directly from the research group leaders. You can find an overview of the Tübingen institutes and clinics and their research priorities on our homepage.

The applications will be reviewed by a PhD Commission, after which some of the candidates will be invited for personal interviews with a selection committee.

The interview

The selection committee consists of three habilitated members of the Faculty of Medicine. The motivation and suitability of the applicant as well as the quality and feasibility of the planned project will be evaluated.

The applicant is expected to give a ten-minute presentation in English (e.g. Powerpoint, maximum 5 slides) about the planned research project. The presentation should include a brief overview of the academic education to date (studies, previous scientific activities), the scientific background of the question, objectives of the research project, a brief description of the methods used and a timetable.

The presentation will be followed by a 15-minute discussion in which the suitability of the candidate and the project itself will be questioned in detail.

selection criteria

• Suitability of the project
• Final grade for previous degrees
• Previous scientific activities
• Interview & Presentation

The multidisciplinary training within the PhD program is offered by representatives and experienced teaching staff of the research focus areas of the Faculty of Medicine Tübingen. The elective program includes innovative techniques like problem based learning courses, case studies and lab rotations. Training is further strengthened by taking advantage of the diverse networks between the research areas and access to state of the art research facilities.

Eligible Modules (with Module Coordinators)

Infection medicine and microbiology.

Prof. Samuel Wagner, PhD

Prodekan, Sektionsleiter

Phone number: 07071 29-84238

E-mail address: [email protected]

Publications: Publications

Personenprofil: Mehr zur Person

Dr. rer. nat. Jana Held

Arbeitsgruppenleiterin

Phone number: 07071 29-85569

Fax number: 07071 29-4684

E-mail address: [email protected]

apl. Prof. Dr. Sandra Beer-Hammer

Stellvertretende Abteilungsleiterin

Phone number: +49 7071 29-74594

Fax number: 07071 29-4942

E-mail address: [email protected]

Professor Alexander Weber, PhD

W3/Full Professor of Innate Immunology

Institution: Institute of Immunology

Address: University of Tübingen

E-mail address: [email protected]

Univ.-Prof. Dr. med. Christian Schürch, MD, PhD

Phone number: +49 7071 29-80201

Phone number: +49 7071 29-2991

E-mail address: [email protected]

Prof. Dr. Lukas Flatz

Oberarzt, Sektionsleiter Dermato-Onkologie

Phone number:

E-mail address: [email protected]

Neurosciences

Marlies Knipper

Professor of Molecular Physiology of Hearing

Institution: Tübingen Hearing Research Center

Address: University Hospital Tübingen

E-mail address: [email protected]

Marius Ueffing

Professor of Ophtalmic Research

Institution: Centre for Ophtalmology

E-mail address: [email protected]

Cardiology, Pharmacology and Diabetology

Prof. Dr. Dr. Bernd Nürnberg

Ärztlicher Direktor der Abteilung

Phone number: 07071 29-74590

E-mail address: [email protected]

Prof. Dr. Reiner Jumpertz-von Schwartzenberg

Leitung Studienzentrale

Phone number: +49 7071 29-68934

E-mail address: [email protected]

Univ.-Prof. Dr. med. Oliver Borst

Ltd. Oberarzt, Heisenberg-Professor Thrombokardiologie

E-mail address: [email protected]

Imaging Science

Univ. Prof. Dr. med. Christian la Fougère

Ärztlicher Direktor Nuklearmedizin und Klinische Molekulare Bildgebung

Phone number: +49 7071 29-86553

Fax number: 07071 29-4601

E-mail address: [email protected]

Bettina Weigelin

Professor of Imaging Science

E-mail address: [email protected]

Biomedical Engineering

Prof. Dr. Andreas Nüssler

Direktor des Siegfried Weller Institut

E-mail address: [email protected]

Dr.Dr.med. Reka Haraszti

Nachwuchsgruppenleiterin und Ärztin in Weiterbildung

Phone number: +49 7071 29-68948

E-mail address: [email protected]

Prof. Dr. rer. nat. Ellen Reisinger

Universitätsprofessorin

Phone number: +49 7071 29-88184

E-mail address: [email protected]

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• Published: 15 May 2024

How Iranian students can master integration into German academia

• Ata Makarem   ORCID: orcid.org/0000-0003-1291-4312 1 , 2 &
• Karel Douglas Klika   ORCID: orcid.org/0000-0001-7009-1487 3  

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Thousands of Iranians study at German universities every year, but many struggle with the German academic system. Here, we offer some advice.

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Knerr, B. International Labor Migration: Asian students in Germany Ch. 6 (Kassel University Press GmbH, 2015).

Hüther, O. & Krücken, G. Higher Education in Germany — Recent Developments in an International Perspective , Vol. 49 (Springer International Publishing, 2018).

Moradi, S. Nat. Hum. Behav. 3 , 1025–1025 (2019).

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MD, PhD Graduates Urged to Become Change Makers

With so much unrest around the world, natural disasters and medical challenges on a global scale, “We hope that you will become leaders in solving these issues,” said Antman.

The May 16 convocation at BU’s Track & Tennis Center bestowed 35 PhD, 144 MD, four combined MD/PhDs, three combined MD/MBAs, one MD/JD and two MD/OMFS (Oral Maxillofacial Surgery) degrees. Fifteen students earned cum laude honors, five magna cum laude and two students, Jonathan Berlowitz and Sarah Golden, graduated summa cum laude .

“The faculty know that you will use the knowledge, the research and clinical skills that you have mastered here to make a difference in the world going forward,” said C. James McKnight , PhD, associate dean and provost of Graduate Medical Sciences.

Two student speakers were chosen by their peers – Josiane Fofana was selected by PhD students.

“(I am) truly humbled, because humility reminds me of how far I’ve come and how much more there is to accomplish,” said Fofana who grew up in Senegal, moving to Boston in 2011 where she completed an associate degree in biological sciences at Bunker Hill Community College, followed by a BS in biochemistry from Brandeis University and finally her PhD in virology, microbiology & immunology at BU.

She asked students to look beyond their degree.

“In the pursuit of knowledge, we often overlook the importance of emotional intelligence,” said Fofana. “Brilliance devoid of empathy just renders us empty, contributing to the injustice in this world.”

Fofana is the mother of a 3-year-old and founded a non-profit providing quality and STEM-based education to children in Dakar, Senegal. She is pursuing a postdoctoral position at the University of Ghana as a Fogarty Global Health Fellow.

“Do not think that the degree or leadership position you’re holding grants you the ultimate wisdom on every issue,” Fofana cautioned. “Remain open to others’ experiences…embrace discomfort in order to grow.”

“One of the most beautiful aspects of our journey together has been the friendships we’ve formed and the shared experiences that have strengthened our bond,” said Merriman. “From our first days of orientation to the challenges of clinical rotations, we’ve grown together, supporting each other every step of the way.”

“I know…that you have the minds, the hearts and the souls of change makers,” she said.

“Healthcare is a basic human right, not a privilege,” she said. “Our profession should focus on improving health beyond the healthcare system – in communities – and focus on prevention of disease with equal focus as treatment of disease,” said Bharel.

At the level of patient interactions, Bharel told students to turn away from their screens, look their patients in the eyes and allow them to tell their story.

“Choose kindness and selflessness. Listen to your patients with humility. Connect your scientific endeavors to our most pressing health issues,” she said.

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Meet Yale Internal Medicine: Allison Gaffey, PhD

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As a part of our “Meet Yale Internal Medicine” series, today’s feature is on Allison Gaffey, PhD, assistant professor (cardiovascular medicine).

As an undergraduate at the University of Connecticut, Allison Gaffey, PhD , assistant professor (cardiovascular medicine), first became interested in reciprocal mind, body, and biological behavioral connections. That interest led her to pursue a career in research – first as a research assistant for studies of health behavior in survivors of physical and psychological trauma, then as a research fellow at the National Institutes of Health (NIH) focused on bio-behavioral interactions within the scope of risk of cancer, and later at Harvard, where she supported investigations of psychosocial stress, emotion, and related autonomic nervous system and endocrine activity.

Altogether these experiences honed her appreciation for how stress and related social circumstances beget poor health, as well as which factors may be protective, and led her to pursue doctoral studies at the University of Notre Dame.

“A PhD in clinical psychology was the ideal way to investigate the effects of stress on related physiology and psychological health, and to also maintain the patient's perspective,” said Gaffey.

Beginning in residency at the Alpert School of Medicine at Brown University, Gaffey says her choice to pursue this work within cardiovascular medicine just made sense.

“Cardiology is an ideal setting for pursuing questions about how stress and other adverse aspects of our daily lives and lifetime experiences ‘get under the skin’ to affect our health,” said Gaffey. “Many people don’t think of psychosocial stress as a risk factor for heart disease, but the data are abundant. This includes examinations of cardiovascular reactivity to short-term, mental stress tasks in a research laboratory or scientific evidence bridging chronic stressors from work, relationships, or discrimination with a greater and earlier incidence of cardiovascular diseases. Stress also directly affects our sleep, another lesser-known cardiovascular risk factor. As cardiovascular disease remains the leading cause of death in the U.S., working in this area is immediately impactful.”

Gaffey came to Yale in 2018, where she trained with Matthew Burg, PhD , professor of medicine (cardiovascular medicine), a preeminent expert in cardiovascular behavioral medicine who investigates the effects of mental stress on cardiovascular physiology. At the same time, she completed an Advanced Research Fellowship in Women’s Health through the VA Connecticut Healthcare System in West Haven, where she trained with Sally Haskell, MD, MS, and Cynthia Brandt, MD, MPH.

“It was the perfect opportunity to simultaneously extend my expertise in cardiovascular medicine and women's health, both in the general population and among Veterans more specifically,” said Gaffey. “It has been estimated that our knowledge of cardiovascular disease in women is about 50 years behind the literature concerning men. There’s an immense need to better understand women, their health, and the exposures that are unique to their lived experiences – as called for by President Biden’s recent executive order for women . Doing this work can also help us to learn about cardiovascular disease in men as well.”

Gaffey, who was promoted to assistant professor earlier this year, recently received a K23 grant , a patient-oriented, mentored career development award, from the National Heart, Lung, and Blood Institute (NHLBI) to support research exploring associations between high blood pressure among younger adults, who are beginning to show high blood pressure earlier in the lifespan.

Gaffey’s research will investigate how stress, childhood trauma, discrimination, and other social vulnerabilities, together influence the increase of blood pressure among younger men and women and may predict the onset of hypertension. She will also examine the role of poor sleep in these associations.

“Many studies have looked at specific types of stress or adversity across the lifespan in relation to cardiovascular risk. In this project, we hope to aggregate those adverse experiences under the umbrella of social vulnerability to see if those who experience greater cumulative vulnerability are more likely to show early increases in blood pressure over time. By doing so, we may learn which men and women need earlier or additional support to manage their blood pressure and when to provide those resources,” said Gaffey.

She is working with data from the CARDIA , a nationwide, prospective cohort study that has been in progress for more than 30 years. Gaffey is conducting a secondary analysis of blood pressure data from that study to determine which factors predict early blood pressure elevations amongst women and men, respectively.

“Although men are diagnosed with hypertension earlier in their lifespan, women show a steeper incline in the rate at which blood pressure increases over their lifespan. Besides certain health conditions and complications associated with pregnancy, we really don't know what social factors are important and implicated in that early risk,” said Gaffey.

As she continues her career, Gaffey hopes to model and identify opportunities for earlier patient-centered prevention and how to test and implement those models in the current healthcare system. She recently co-authored a perspective with Burg and Bruce Rollman, MD, MPH (University of Pittsburgh Medical Center), encouraging the American Heart Association to include psychological health – particularly depression and anxiety – as one of their risk factors for cardiovascular disease in the next update to their Life’s Essential 8 heart health metric.

“Depression, post-traumatic stress disorder, trauma, anxiety, life stress, and other social vulnerabilities are associated with risk for cardiovascular disease,” said Gaffey. “In line with the U.S. Preventive Services Task Force recommendations, screening for depression and anxiety alone would be a meaningful first step towards better addressing these risk factors and more holistic, patient-centered care.”

Gaffey tells her trainees that it is important to surround yourself with people who have varied backgrounds and interests, who can offer different types of support while allowing the freedom needed to develop your unique scientific, clinical, educational, and policy-focused interests.

“In academic medicine, it’s vital to recognize the value of your unique expertise and the power of your voice, but also to use your voice to reflect the patient’s perspective – their lived experiences and motivations,” said Gaffey. “There’s fulfillment in giving voice to our patients.”

The Department of Internal Medicine at Yale School of Medicine is among the nation's premier departments, bringing together an elite cadre of clinicians, investigators, educators, and staff in one of the world's top medical schools. To learn more, visit Internal Medicine.

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30 years of (re)productive biology: a symposium honoring professor Martin M. Matzuk, MD, PhD

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T Rajendra Kumar, Kathleen H Burns, Wei Yan, 30 years of (re)productive biology: a symposium honoring professor Martin M. Matzuk, MD, PhD, Biology of Reproduction , Volume 110, Issue 5, May 2024, Pages 852–853, https://doi.org/10.1093/biolre/ioae051

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Martin Matzuk, MD, PhD, is a professor and the Stuart A. Wallace Chair of the Department of Pathology and Immunology at the Baylor College of Medicine, Houston, TX, USA. During the span of over 30 years at a single institution, he has made seminal contributions to various areas of reproductive biology research, including systematically deciphering the developmental roles of TGF-β growth factor family members (inhibins, activins, activin receptor-2, follistatin, GDF9, BMPs, and SMADs) in normal physiology and pathology of ovarian, testicular, and uterine functions. In the last decade, he developed a comprehensive drug development center at Baylor College of Medicine embarking on rapid target identification using DNA-encoded chemistry and in silico screening. In addition to continuing basic research, his team is now mostly focusing on the development of highly specific drugs for male contraception, endometriosis and ovarian cancer, and many other diseases.

Dr Matzuk received numerous honors and awards, including the Elected Member of the National Academy of Sciences, the Carl Hartman Award, the highest award conferred by the SSR, and, more recently, The Texas State Hill Prize. He trained numerous undergraduate, graduate, clinical, and postdoctoral fellows who have made distinguished careers in basic science and clinical medicine. Many of his former trainees and some of his collaborators met in person in April 2023 and felicitated his achievements in a day-long scientific symposium organized by the three of us (TRK, KHB, and WY) at Baylor College of Medicine, Houston, TX ( Figure 1).

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Jewish doctors launch group to combat antisemitism in medicine: ‘it’s nazi germany all over again’.

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Jewish doctors alarmed by “rampant” antisemitism and even violence at health-care institutions in the US have launched their own national advocacy group to combat the hatred in medicine. 

Thousands of medical professionals from across the country have joined the new American Jewish Medical Association to stand up for Jews, the group recently told The Post.

“It’s fundamentally scary for those of us who care about humanity. It’s Nazi Germany all over again,” said Manhattan plastic surgeon Yael Halaas, founder and president of the new group and a graduate of Columbia University and Cornell Medical School.

The hatred leveled against Jews after the Palestinian terror group Hamas’ Oct. 7 invasion of Israel hit home for Halaas, whose Jewish parents emigrated to the US from Cuba and Argentina.

Many members of her dad’s family were killed in Auschwitz during the Holocaust.

Medical schools particularly have become hostile environments for Jewish students, the organizers said.

Among the troubling incidents:

-George Washington University’s medical school hosted a faculty panel in December that declared the terrorists have a “right of resistance” against Israel.

-Some medical professionals have praised the Hamas Oct. 7 attack on Israel and denied the fact that victims in Israel were sexually assaulted.

— A sign that read, “Free Palestine from Nazi Zionist Schwein” — the German word meaning “pig” — appeared in front of the cancer center at the University of California at San Francisco’s medical school.

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—  Anti-Israel protesters at rallies at Columbia University’s Medical School/Mailman School of Public Health have chanted “From the River to the Sea,” a phrase many interpret as calling for the elimination of Israel and celebrating suicide bombers who kill Israelis.

“The Jewish medical students are bullied into silence,” Halaas said. “They are being ostracized. It shouldn’t be controversial to say Israel is integral to Jewish identity.

“Anti-Zionism is antisemitism,” she said.

The term “zionism” refers to the right of the Jewish state to exist.

Orthopedist Cary Schwartzbach, a Queens native who practices in Fairfax, Va., and a son of Holocaust survivors and treasurer of the new group, said, “Oct. 7 was a wake-up call.

“It opened the gates for antisemitism,” the doctor said.

“There’s a sense of hostilities against Jews. Antisemitism has made a lot of hospital residency programs uncomfortable for Jewish students,” he said.

“We need a voice. We need to protect ourselves. We need to protect medical school students.”

Larisa Geskin, an oncologist and director of the Comprehensive Skin Cancer Center at Columbia University’s Medical Center who emigrated from Latvia in the former Soviet Union when she was 20, said she was stunned by the antisemitism she’s personally witnessed on campus.

“There is rampant anti semitism in the medical community,” the 56-year-old doctor said.

“This is so familiar to me. In the Soviet Union antisemitism was rampant. I never thought I’d hear the same rhetoric in the United States that I heard in the Soviet Union,” she said.

Given how Jews have been abused throughout history, the doctors said it’s horrifying to hear propaganda painting Israelis as white oppressors.

“Jews are also a minority. We need protection like other minorities,” Geskin said.

“We see history repeating itself. We feel we have to say something. They were chanting `From the River to the Sea’ in my backyard. This is what is happening in the medical community.”

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• http://orcid.org/0000-0002-7127-6560 Fábio Carlos Lucas de Oliveira 1 , 2
• 1 École de kinésiologie et de loisir , Université de Moncton , Moncton , NB , Canada
• 2 Research Unit in Sport and Physical Activity (CIDAF) , University of Coimbra Faculty of Sport Sciences and Physical Education , Coimbra , Portugal
• Correspondence to Dr Fábio Carlos Lucas de Oliveira, Université de Moncton, Moncton, NB, Canada; fclud{at}ulaval.ca

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• Athletic Tape

What did I do?

My PhD project aimed to determine whether kinesiotaping provided additional benefits for treating individuals with rotator cuff-related shoulder pain (RCRSP). To achieve this goal, I investigated (1) the alterations in the rotator cuff (RC) muscle activation, (2) the immediate effects of kinesiotaping on the acromiohumeral distance (AHD) and shoulder proprioception and (3) the effects of kinesiotaping on the symptoms, functional limitations and underlying deficits associated with RCRSP.

Why did I do it?

The application of kinesiotaping has been a popular approach in clinics. However, few studies have investigated its immediate, short-term or isolated effects among individuals with RCRSP. 4 Current evidence has been insufficient to conclude the effectiveness of kinesiotaping in improving pain and function in this population. 5 Therefore, there was a need for high-quality evidence to better guide clinicians on whether to use kinesiotaping for the rehabilitation of individuals with RCRSP.

How did I do it?

First, I conducted a systematic review 6 to synthesise the evidence concerning the electromyographic activation of RC muscles in individuals with RCRSP. This study guided which RC and scapulothoracic muscles should be targeted for treatment and how these muscles should be rehabilitated. Based on this work, I designed a 6-week exercise-based rehabilitation programme as the cornerstone of my PhD project.

To assess underlying deficits, I conducted a cross-sectional study 7 that investigated whether kinesiotaping provides an immediate increase in the AHD (assessed using an ultrasound scanner) and whether the active joint repositioning sense (assessed using inertial measurements unit wireless sensors) was changed in individuals with RCRSP.

Lastly, I conducted a randomised controlled trial 4 (RCT) to identify whether adding kinesiotaping to the treatments would provide a faster and impactful difference to the healing of individuals with RCRSP. Fifty-two (52) individuals with RCRSP were recruited. Participants had at least one positive sign in each category of the following clinical tests 8 : (a) painful arc of movement (flexion and abduction); (b) Neer or Hawkins-Kennedy impingement test and (c) pain during resisted external rotation, abduction or empty can test.

Primary outcome (functional limitations) was assessed using the Disabilities of the Arm (DASH) questionnaire at baseline, week 3, week 6, week 12 and 6 months. Secondary outcomes were evaluated at commencement (baseline) and following the treatment (week 6), which included: pain intensity assessed using the Brief Pain Intensity (BPI), symptoms assessed using the Western Ontario Rotator Cuff (WORC) index, pain-free and full range of motion (ROM) (shoulder flexion and abduction) assessed using a universal goniometer, and AHD at rest and 60° of shoulder abduction assessed using an ultrasound scanner.

All participants were evaluated at baseline and randomly divided into experimental or control groups. Both groups received a 6-week rehabilitation programme composed of 10 physiotherapy sessions. Kinesiotaping was added to the treatment of individuals allocated in the experimental group ( figure 1 ). Halfway through treatment (week 3), self-report outcomes (DASH, BPI and WORC) were re-evaluated to assess functional limitations and symptoms. At the end of treatment (week 6), all outcomes were re-evaluated as performed at baseline. Participants received four exercises tailored to individual needs to be performed at home throughout the treatment and during follow-up periods. Self-report outcomes were also re-evaluated 12 weeks and 6 months after the first physiotherapy session to identify continued changes.

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Kinesiotaping technique for rotator cuff-related shoulder pain and underlying deficits. First strip ((1) Y-shape surrounding deltoid muscles), second strip ((2) I-shape in functional correction for multiaxial shoulder instability over the glenohumeral joint, supraspinatus, trapezius and middle deltoid muscles) and third strip ((3) I-shape in mechanical correction for glenohumeral joint).

What did I find?

The systematic review 6 revealed strong evidence that the infraspinatus and supraspinatus muscle activities were not altered during isometric contractions. Moderate evidence indicated that alterations in the RC muscle activities are best observed during dynamic tasks.

The cross-sectional study showed that kinesiotaping induced an increase (p<0.001) in AHD at 60° abduction in individuals with RCRSP, whereas no immediate effect was observed with the proprioception.

The RCT 4 demonstrated that functional limitations, symptoms, ROM and AHD improved in both groups, although adding kinesiotaping did not lead to superior outcomes compared with the exercise-based treatment alone in the mid-term and long term for individuals with RCRSP.

What is the most important clinical impact/practical application?

This PhD project revealed that kinesiotaping is not an essential element for rehabilitating individuals with RCRSP. Individuals who used kinesiotaping did not present with faster or more impactful results than those who did not, evidencing that kinesiotaping did not provide additional benefits for treating individuals with RCRSP in the mid-term and long term. Subsequently, clinicians and healthcare professionals should not expect supplementary gains from kinesiotaping when added to a rehabilitation programme based on sensorimotor exercises for individuals with RCRSP.

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This study involves human participants and protocol was registered at ClinicalTrials.gov ( NCT02881021 ) on 25 August 2016. Ethics approval was obtained from the Ethics Committee of Quebec Rehabilitation Institute (IRDPQ #2016-496) of the Centre Integrated University of Health and Social Services (CIUSSS-CN). Participants gave informed consent to participate in the study before taking part.

• Maenhout AG ,
• Vanderstukken F , et al
• Hébert LJ , et al
• van der Heijden GJ
• de Oliveira FCL ,
• Pairot de Fontenay B ,
• Bouyer LJ , et al
• Saracoglu I ,
• Bouyer LJ ,
• Ager AL , et al
• Michener LA ,
• Walsworth MK ,
• Doukas WC , et al

X @fabiophysio

Contributors FCLO contributed to conception, design and preparation of the procedures. He also conducted the recruitment of participants, data collection, the rehabilitation programme, analyses and data interpretation, and writing.

Funding This work was supported by the Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (CAPES). FCLO received a doctoral scholarship from the Brazilian Government through the Science without Borders programme in association with the CAPES Foundation.

Competing interests None declared.

Provenance and peer review Not commissioned; externally peer reviewed.

Author note This doctoral project was conducted at the Centre interdisciplinaire de recherche en réadaptation et intégration sociale (Cirris), Université Laval, Québec City, Canada.

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Monoallelic TYROBP deletion is a novel risk factor for Alzheimer’s disease

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Biallelic loss-of-function variants in TYROBP and TREM2 cause autosomal recessive presenile dementia with bone cysts known as Nasu-Hakola disease (NHD, alternatively polycystic lipomembranous osteodysplasia with sclerosing leukoencephalopathy, PLOSL). Some other TREM2 variants contribute to the risk of Alzheimer’s disease (AD) and frontotemporal dementia, while deleterious TYROBP variants are globally extremely rare and their role in neurodegenerative diseases remains unclear. The population history of Finns has favored the enrichment of deleterious founder mutations, including a 5.2 kb deletion encompassing exons 1-4 of TYROBP and causing NHD in homozygous carriers. We used here a proxy marker to identify monoallelic TYROBP deletion carriers in the Finnish biobank study FinnGen combining genome and health registry data of 520,210 Finns. We show that monoallelic TYROBP deletion associates with an increased risk and earlier onset age of AD and dementia when compared to noncarriers. In addition, we present the first reported case of a monoallelic TYROBP deletion carrier with NHD-type bone cysts. Mechanistically, monoallelic TYROBP deletion leads to decreased levels of DAP12 protein (encoded by TYROBP ) in myeloid cells. Using transcriptomic and proteomic analyses of human monocyte-derived microglia-like cells, we show that upon lipopolysaccharide stimulation monoallelic TYROBP deletion leads to the upregulation of the inflammatory response and downregulation of the unfolded protein response when compared to cells with two functional copies of TYROBP . Collectively, our findings indicate TYROBP deletion as a novel risk factor for AD and suggest specific pathways for therapeutic targeting.

One Sentence Summary Nasu-Hakola disease causing TYROBP deletion increases the risk of Alzheimer’s disease in elderly monoallelic carriers in the Finnish population.

Competing Interest Statement

CH collaborates with Denali Therapeutics and is a member of the advisory boards of AviadoBio and Cure Ventures. The other authors declare that they have no competing interests.

Funding Statement

This study was funded by the following: Research Council of Finland (to HM, MH, MT, VL). Faculty of Health Sciences, University of Eastern Finland (to HM, MT). Sigrid Juselius Foundation (to MH, VL, ES, TN). The Strategic Neuroscience Funding of the University of Eastern Finland (to MH, AH, VL, ES). Alzheimer's Association (to MH). The State Research Funding KUH-VTR (to VL, ES). Doctoral Programme in Molecular Medicine (to RMW, HJ). Deutsche Forschungsgemeinschaft (DFG, German Research Foundation) under Germany's Excellence Strategy within the framework of the Munich Cluster for Systems Nuerology, SyNergy (to CH and SFL) and a Koselleck Project (to CH). JPco-fuND 2019 Personalised Medicine for Neurodegenerative Diseases; PMG-AD (to CH, SFL, JCL, and MH).

Author Declarations

I confirm all relevant ethical guidelines have been followed, and any necessary IRB and/or ethics committee approvals have been obtained.

The details of the IRB/oversight body that provided approval or exemption for the research described are given below:

Medical Research Ethics committee of Wellbeing Services County of North Savo gave ethical approval for this work.

I confirm that all necessary patient/participant consent has been obtained and the appropriate institutional forms have been archived, and that any patient/participant/sample identifiers included were not known to anyone (e.g., hospital staff, patients or participants themselves) outside the research group so cannot be used to identify individuals.

I understand that all clinical trials and any other prospective interventional studies must be registered with an ICMJE-approved registry, such as ClinicalTrials.gov. I confirm that any such study reported in the manuscript has been registered and the trial registration ID is provided (note: if posting a prospective study registered retrospectively, please provide a statement in the trial ID field explaining why the study was not registered in advance).

I have followed all appropriate research reporting guidelines, such as any relevant EQUATOR Network research reporting checklist(s) and other pertinent material, if applicable.

Data Availability

All data associated with this study are present in the paper or the Supplementary Materials. The study participant consent does not allow opening the sequencing (WGS, RNA-seq) or proteomic data generated and analyzed during the current study, but they are available from the corresponding authors (H.M. or M.H.) on reasonable request. Summary statistics from each FinnGen data release will be made publicly available after a one-year embargo period and can be accessed freely at www.finngen.fi/en/access_results. For individual level data, the Finnish biobank data can be accessed through the Fingenious services (https://site.fingenious.fi/en/) managed by FINBB. Access to Finnish Health register data can be applied from Findata (https://findata.fi/en/data/).

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Developmental Biology

Vishnu Saraswathy, PhD., and Yuxiao Xu named 2024 Hope Center Award Winners

Congratulations to Vishnu Saraswathy and Yuxiao Xu!

Vishnu Saraswathy, Posdoc in Mayssa Mokalled’s Lab, and Yuxiao Xu, Graduate Research Assistant in Mayssa Mokalled’s Lab, have been named as two of the 2024 Hope Center Awards winners